Project 793: R. J. Garwood, J. A. Dunlop, G. Giribet, M. D. Sutton. 2011. Anatomically modern Carboniferous harvestmen demonstrate early cladogenesis and stasis in Opiliones. Nature Communications. 2 (1):null.
Abstract
Harvestmen, the third most-diverse arachnid order, are an ancient group found on all continental landmasses, except Antarctica. However, a terrestrial mode of life and leathery, poorly mineralized exoskeleton makes preservation unlikely, and their fossil record is limited. The few Palaeozoic species discovered to date appear surprisingly modern, but are too poorly preserved to allow unequivocal taxonomic placement. Here, we use high-resolution X-ray micro-tomography to describe two new harvestmen from the Carboniferous (~305 Myr) of France. The resulting computer models allow the first phylogenetic analysis of any Palaeozoic Opiliones, explicitly resolving both specimens as members of different extant lineages, and providing corroboration for molecular estimates of an early Palaeozoic radiation within the order. Furthermore, remarkable similarities between these fossils and extant harvestmen implies extensive morphological stasis in the order. Compared with other arachnids—and terrestrial arthropods generally—harvestmen are amongst the first groups to evolve fully modern body plans. This project builds upon MorphoBank Project 631, but includes new fossils.Read the article »
Article DOI: 10.1038/ncomms1458
Project DOI: 10.7934/P793, http://dx.doi.org/10.7934/P793
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 9850 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 253 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 542 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:253/0 |
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MorphoBank Project 793
MorphoBank Project 793
- Creation Date:
19 November 2011 - Publication Date:
14 January 2012 - Project views: 46231
- Matrix downloads: 13
Authors' Institutions
- Museum für Naturkunde - Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin
- Harvard University
- Imperial College London
- University of Manchester
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||||
Gonzalo Giribet Project Administrator | 7 | 0 | 1 | 0 | 0 | 0 | 0 | 1036 (610, 327, 99) | 0 | 0 | 0 | |||||
MorphoBank Curator Full membership | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | |||||
Jason Dunlop Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | |||||
Russell Garwood Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | |||||
Prashant Sharma Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | |||||
Jo Wolfe Full membership | 44 | 0 | 0 | 253 | 0 | 0 | 0 | 8814 (7617, 0, 1197) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M1802' (51 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Limulus polyphemus Last Modified in 12/05/13 | 42 | 161 | 161 | 0 | 50 | 0 | 0 | 6 |
[2] Carcinoscorpius rotundicauda Last Modified in 12/05/13 | 44 | 159 | 159 | 0 | 50 | 0 | 0 | 6 |
[3] Androctonus australis Last Modified in 12/05/13 | 32 | 176 | 176 | 0 | 45 | 0 | 0 | 6 |
[4] Belisarius xambeui Last Modified in 12/05/13 | 38 | 166 | 166 | 0 | 49 | 0 | 0 | 6 |
[5] Siro rubens Last Modified in 12/05/13 | 33 | 189 | 189 | 1 | 30 | 0 | 0 | 6 |
[6] Parasiro coiffaiti Last Modified in 12/05/13 | 48 | 175 | 175 | 0 | 30 | 0 | 0 | 6 |
[7] Chileogovea oedipus Last Modified in 12/05/13 | 140 | 98 | 98 | 1 | 14 | 0 | 0 | 6 |
[8] Ogovea cameroonensis Last Modified in 12/05/13 | 141 | 97 | 97 | 0 | 15 | 0 | 0 | 6 |
[9] Huitaca ventralis Last Modified in 12/05/13 | 141 | 96 | 96 | 0 | 16 | 0 | 0 | 6 |
[10] Meghalaya sp. 19a Last Modified in 12/05/13 | 129 | 108 | 107 | 2 | 14 | 0 | 0 | 6 |
[11] Fangensis insulanus Last Modified in 12/05/13 | 47 | 178 | 178 | 1 | 27 | 0 | 0 | 6 |
[12] Leptopsalis sp. 3 gunung bondeh Last Modified in 12/05/13 | 48 | 177 | 177 | 1 | 27 | 0 | 0 | 6 |
[13] Dalquestia formosa Last Modified in 12/05/13 | 48 | 186 | 186 | 0 | 19 | 0 | 0 | 6 |
[14] Odiellus troguloides Last Modified in 12/05/13 | 35 | 196 | 196 | 0 | 22 | 0 | 0 | 6 |
[15] Phalangium opilio Last Modified in 12/05/13 | 34 | 197 | 197 | 0 | 22 | 0 | 0 | 6 |
[16] Opilio parietinus Last Modified in 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 6 |
[17] Astrobunus grallator Last Modified in 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 6 |
[18] Nelima silvatica Last Modified in 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 6 |
[19] Leiobunum sp. Last Modified in 12/05/13 | 32 | 199 | 199 | 0 | 22 | 0 | 0 | 6 |
[20] Hadrobunus maculosus Last Modified in 12/05/13 | 47 | 187 | 187 | 0 | 19 | 0 | 0 | 6 |
[21] Caddo agilis Last Modified in 12/05/13 | 48 | 186 | 186 | 0 | 19 | 0 | 0 | 6 |
[22] Ischyropsalis luteipes Last Modified in 12/05/13 | 35 | 191 | 191 | 0 | 27 | 0 | 0 | 6 |
[23] Hesperonemastoma modestum Last Modified in 12/05/13 | 47 | 180 | 180 | 0 | 26 | 0 | 0 | 6 |
[24] Ceratolasma tricantha Last Modified in 12/05/13 | 48 | 181 | 181 | 0 | 24 | 0 | 0 | 6 |
[25] Sabacon cavicolens Last Modified in 12/05/13 | 47 | 182 | 182 | 0 | 24 | 0 | 0 | 6 |
[26] Taracus sp. Last Modified in 12/05/13 | 47 | 182 | 182 | 0 | 24 | 0 | 0 | 6 |
[27] Dicranolasma soerenseni Last Modified in 12/05/13 | 47 | 180 | 180 | 0 | 26 | 0 | 0 | 6 |
[28] Centetostoma dubium Last Modified in 12/05/13 | 50 | 177 | 177 | 0 | 26 | 0 | 0 | 6 |
[29] Ortholasma sp. Last Modified in 12/05/13 | 48 | 180 | 180 | 0 | 25 | 0 | 0 | 6 |
[30] Nemastoma bimaculatum Last Modified in 12/05/13 | 33 | 191 | 191 | 0 | 29 | 0 | 0 | 6 |
[31] Nipponopsalis abei Last Modified in 12/05/13 | 48 | 180 | 180 | 0 | 25 | 0 | 0 | 6 |
[32] Trogulus nepaeformis Last Modified in 12/05/13 | 34 | 190 | 190 | 0 | 29 | 0 | 0 | 6 |
[33] † Halitherses grimaldii Last Modified in 12/05/13 | 129 | 86 | 86 | 18 | 20 | 0 | 0 | 6 |
[34] † Ameticos scolos Last Modified in 12/05/13 | 145 | 58 | 58 | 33 | 17 | 0 | 0 | 6 |
[35] † Macrogyion cronus Last Modified in 12/05/13 | 115 | 64 | 64 | 59 | 15 | 0 | 0 | 6 |
[36] † Eophalangium sheari Last Modified in 12/05/13 | 116 | 51 | 51 | 75 | 11 | 0 | 0 | 6 |
[37] † Hastocularis argus Last Modified in 12/05/13 | 104 | 72 | 71 | 71 | 7 | 0 | 0 | 6 |
[38] † Mesobunus dunlopi Last Modified in 12/05/13 | 121 | 60 | 60 | 65 | 7 | 0 | 0 | 6 |
[39] Equitius doriae Last Modified in 12/05/13 | 56 | 175 | 175 | 0 | 22 | 0 | 0 | 6 |
[40] Triaenobunus sp. Last Modified in 12/05/13 | 58 | 173 | 173 | 0 | 22 | 0 | 0 | 6 |
[41] Zuma acuta Last Modified in 12/05/13 | 58 | 173 | 173 | 0 | 22 | 0 | 0 | 6 |
[42] Sandokan cf. alticeps Last Modified in 12/05/13 | 47 | 184 | 183 | 0 | 23 | 0 | 0 | 6 |
[43] Gnomulus sp. Last Modified in 12/05/13 | 47 | 184 | 183 | 0 | 23 | 0 | 0 | 6 |
[44] Scotolemon lespesi Last Modified in 12/05/13 | 35 | 192 | 192 | 0 | 26 | 0 | 0 | 6 |
[45] Maiorerus randoi Last Modified in 12/05/13 | 38 | 187 | 187 | 0 | 28 | 0 | 0 | 6 |
[46] Bishopella laciniosa Last Modified in 12/05/13 | 45 | 183 | 183 | 0 | 25 | 0 | 0 | 6 |
[47] Gnidia holmbergi Last Modified in 12/05/13 | 43 | 185 | 185 | 0 | 25 | 0 | 0 | 6 |
[48] Pachyloides thorellii Last Modified in 12/05/13 | 34 | 193 | 193 | 0 | 26 | 0 | 0 | 6 |
[49] Hoplobunus sp. Last Modified in 12/05/13 | 45 | 181 | 181 | 0 | 27 | 0 | 0 | 6 |
[50] Solifugae Last Modified in 12/05/13 | 34 | 157 | 157 | 0 | 62 | 0 | 0 | 6 |
[51] Pseudoscorpiones Last Modified in 12/05/13 | 34 | 163 | 163 | 0 | 56 | 0 | 0 | 6 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 46231 | |
Project overview | 3744 | |
Media views | 7985 | Media search (5409 views); M329570 (858 views); M149206 (825 views); M149204 (893 views); |
Taxon list | 20672 | |
Matrix views | 2803 | Matrix landing page (2448 views); Anatomically modern Carboniferous harvestmen (355 views); |
Bibliography | 2013 | |
Specimen list | 5202 | |
Documents list | 1974 | |
Views for media list | 1838 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 333 | |
Matrix downloads | 13 | Anatomically modern Carboniferous harvestmen (13 downloads); |
Project downloads | 319 | |
Document downloads | 1 | Copyright Permission for Image (1 download); |