Project 4400: I. Yáñez, D. Pol, J. Leardi, O. Alcober, R. Martinez. 2021. An enigmatic new archosauriform from the Carnian-Norian Ischigualasto Formation of Northwestern Argentina. Acta Palaeontologica Polonica. 66 (3):509-533.
Abstract
In this contribution we introduce a new Late Triassic archosaur, Incertovenator longicollum gen. et sp. nov., with an unusual combination of character states that are present in certain early avemetatarsalian and pseudosuchian archosaur clades. The holotype consists of a partial postcranial skeleton, preserving most of the axial skeleton and displaying a marked anteroposterior elongation in the cervical vertebrae. We include I. longicollum gen. et sp. nov. into one of the most comprehensive early archosaur phylogenetic data sets available, and recover it as either an early diverging avemetatarsalian, closely associated with the clade Aphanosauria and Ornithodira, or as an early diverging loricatan closely related to Mandasuchus tanyauchen in the most parsimonious trees. We further evaluate which alternative phylogenetic positions can I. longicollum gen. et sp. nov. take in the suboptimal trees, and determined which character states support those alternative positions in comparison with those of the unconstrained analysis. The analyses recover the new taxon in three main general phylogenetic placements within Archosauria, as well as one position outside this clade, highlighting widespread morphological evolutionary convergence towards neck elongation in several clades of Triassic archosauriforms.Read the article »
Article DOI: 10.4202/app.00806.2020
Project DOI: 10.7934/P4400, http://dx.doi.org/10.7934/P4400
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 22907 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 419 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 916 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:400/19 |
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MorphoBank Project 4400
MorphoBank Project 4400
- Creation Date:
12 September 2022 - Publication Date:
31 October 2022 - Project views: 6064
- Matrix downloads: 3
Authors' Institutions
- Museo Paleontológico Egidio Feruglio
- Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)
- Universidad de Buenos Aires
- Universidad Nacional de San Juan
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| MorphoBank Curator Project Administrator | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| MorphoBank Curator Full membership | 99 | 1 | 1 | 419 | 0 | 0 | 22907 (22815, 0, 92) | 0 | 0 | 0 | ||||
| Imanol Yáñez Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M29022' (99 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] † Mesosuchus browni Last Modified in 10/31/22 | 84 | 335 | 335 | 0 | 0 | 0 | 0 | 3 |
| [2] † Prolacerta broomi Last Modified in 10/31/22 | 56 | 363 | 363 | 0 | 0 | 0 | 0 | 3 |
| [3] † Archosaurus rossicus Last Modified in 10/31/22 | 414 | 5 | 5 | 0 | 0 | 0 | 0 | 3 |
| [4] † Proterosuchus Last Modified in 10/31/22 | 51 | 368 | 368 | 0 | 0 | 0 | 0 | 3 |
| [5] † Erythrosuchus africanus Last Modified in 10/31/22 | 50 | 369 | 369 | 0 | 0 | 0 | 0 | 3 |
| [6] † Vancleavea campi Last Modified in 10/31/22 | 134 | 285 | 285 | 0 | 0 | 0 | 0 | 3 |
| [7] † Chanaresuchus bonapartei Last Modified in 10/31/22 | 53 | 367 | 366 | 0 | 0 | 0 | 0 | 3 |
| [8] † Tropidosuchus romeri Last Modified in 10/31/22 | 103 | 317 | 316 | 0 | 0 | 0 | 0 | 3 |
| [9] † Euparkeria capensis Last Modified in 10/31/22 | 18 | 402 | 401 | 0 | 0 | 0 | 0 | 3 |
| [10] † Parasuchus hislopi Last Modified in 10/31/22 | 66 | 353 | 353 | 0 | 0 | 0 | 0 | 3 |
| [11] † Smilosuchus gregorii Last Modified in 10/31/22 | 76 | 343 | 343 | 0 | 0 | 0 | 0 | 3 |
| [12] † Pseudopalatus pristinus Last Modified in 10/31/22 | 33 | 386 | 386 | 0 | 0 | 0 | 0 | 3 |
| [13] † Gracilisuchus stipanicicorum Last Modified in 10/31/22 | 108 | 315 | 310 | 0 | 1 | 0 | 0 | 3 |
| [14] † Turfanosuchus dabanensis Last Modified in 10/31/22 | 147 | 273 | 272 | 0 | 0 | 0 | 0 | 3 |
| [15] † Yonghesuchus sangbiensis Last Modified in 10/31/22 | 328 | 91 | 90 | 0 | 1 | 0 | 0 | 3 |
| [16] † Incertovenator longicollum Last Modified in 10/31/22 | 382 | 35 | 35 | 0 | 2 | 0 | 0 | 3 |
| [17] † Ornithosuchus longidens Last Modified in 10/31/22 | 140 | 279 | 279 | 0 | 0 | 0 | 0 | 3 |
| [18] † Riojasuchus tenuisceps Last Modified in 10/31/22 | 62 | 357 | 357 | 0 | 0 | 0 | 0 | 3 |
| [19] † Revueltosaurus callenderi Last Modified in 10/31/22 | 47 | 372 | 372 | 0 | 0 | 0 | 0 | 3 |
| [20] † Stagonolepis robertsoni Last Modified in 10/31/22 | 101 | 318 | 318 | 0 | 0 | 0 | 0 | 3 |
| [21] † Aetosaurus ferratus Last Modified in 10/31/22 | 152 | 268 | 267 | 0 | 0 | 0 | 0 | 3 |
| [22] † Longosuchus meadei Last Modified in 10/31/22 | 70 | 349 | 349 | 0 | 0 | 0 | 0 | 3 |
| [23] † Ticinosuchus ferox Last Modified in 10/31/22 | 211 | 208 | 208 | 0 | 0 | 0 | 0 | 3 |
| [24] † Qianosuchus mixtus Last Modified in 10/31/22 | 131 | 288 | 288 | 0 | 0 | 0 | 0 | 3 |
| [25] † Xilousuchus sapingensis Last Modified in 10/31/22 | 300 | 119 | 119 | 0 | 0 | 0 | 0 | 3 |
| [26] † Arizonasaurus babbitti Last Modified in 10/31/22 | 163 | 256 | 256 | 0 | 0 | 0 | 0 | 3 |
| [27] † Poposaurus gracilis holotype Last Modified in 10/31/22 | 380 | 38 | 38 | 0 | 1 | 0 | 0 | 3 |
| [28] † Poposaurus gracilis yale Last Modified in 10/31/22 | 202 | 217 | 216 | 0 | 1 | 0 | 0 | 3 |
| [29] † Lotosaurus adentus Last Modified in 10/31/22 | 103 | 316 | 316 | 0 | 0 | 0 | 0 | 3 |
| [30] † Sillosuchus longicervix Last Modified in 10/31/22 | 338 | 80 | 80 | 0 | 1 | 0 | 0 | 3 |
| [31] † Effigia okeeffeae Last Modified in 10/31/22 | 98 | 320 | 320 | 0 | 1 | 0 | 0 | 3 |
| [32] † Shuvosaurus inexpectatus Last Modified in 10/31/22 | 107 | 311 | 311 | 0 | 1 | 0 | 0 | 3 |
| [33] † Prestosuchus chiniquensis Last Modified in 10/31/22 | 245 | 174 | 174 | 0 | 0 | 0 | 0 | 3 |
| [34] † UFRGS 0156 T Last Modified in 10/31/22 | 251 | 168 | 168 | 0 | 0 | 0 | 0 | 3 |
| [35] † UFRGS 152 T Last Modified in 10/31/22 | 151 | 268 | 268 | 0 | 0 | 0 | 0 | 3 |
| [36] † Combined prestosuchus Last Modified in 10/31/22 | 61 | 358 | 358 | 0 | 0 | 0 | 0 | 3 |
| [37] † Saurosuchus galilei Last Modified in 10/31/22 | 109 | 310 | 310 | 0 | 0 | 0 | 0 | 3 |
| [38] † Batrachotomus kuperferzellensis Last Modified in 10/31/22 | 63 | 356 | 356 | 0 | 0 | 0 | 0 | 3 |
| [39] † Fasolasuchus tenax Last Modified in 10/31/22 | 256 | 163 | 163 | 0 | 0 | 0 | 0 | 3 |
| [40] † Rauisuchus triradentes Last Modified in 10/31/22 | 285 | 134 | 134 | 0 | 0 | 0 | 0 | 3 |
| [41] † Polonosuchus silesiacus Last Modified in 10/31/22 | 300 | 119 | 119 | 0 | 0 | 0 | 0 | 3 |
| [42] † Postosuchus kirkpatricki Last Modified in 10/31/22 | 76 | 343 | 343 | 0 | 0 | 0 | 0 | 3 |
| [43] † Postosuchus alisonae Last Modified in 10/31/22 | 249 | 170 | 170 | 0 | 0 | 0 | 0 | 3 |
| [44] † CM 73372 Last Modified in 10/31/22 | 255 | 163 | 163 | 0 | 1 | 0 | 0 | 3 |
| [45] † Hesperosuchus agilis AMNH-6758 Last Modified in 10/31/22 | 200 | 219 | 219 | 0 | 0 | 0 | 0 | 3 |
| [46] † Dromicosuchus grallator Last Modified in 10/31/22 | 145 | 274 | 274 | 0 | 0 | 0 | 0 | 3 |
| [47] † Hesperosuchus agilis UCMP-12947 Last Modified in 10/31/22 | 131 | 288 | 288 | 0 | 0 | 0 | 0 | 3 |
| [48] † Dibothrosuchus elaphros Last Modified in 10/31/22 | 198 | 220 | 220 | 0 | 1 | 0 | 0 | 3 |
| [49] † Terrestrisuchus gracilis Last Modified in 10/31/22 | 100 | 318 | 318 | 0 | 1 | 0 | 0 | 3 |
| [50] † Sphenosuchus acutus Last Modified in 10/31/22 | 201 | 218 | 218 | 0 | 0 | 0 | 0 | 3 |
| [51] † Litargosuchus leptorhynchus Last Modified in 10/31/22 | 229 | 190 | 190 | 0 | 0 | 0 | 0 | 3 |
| [52] † Kayentasuchus walkeri Last Modified in 10/31/22 | 310 | 108 | 108 | 0 | 1 | 0 | 0 | 3 |
| [53] † Orthosuchus stormbergi Last Modified in 10/31/22 | 96 | 322 | 322 | 0 | 1 | 0 | 0 | 3 |
| [54] Alligator mississippiensis Last Modified in 10/31/22 | 46 | 374 | 372 | 0 | 1 | 0 | 0 | 3 |
| [55] † Protosuchus haughtoni Last Modified in 10/31/22 | 260 | 159 | 159 | 0 | 0 | 0 | 0 | 3 |
| [56] † Protosuchus richardsoni Last Modified in 10/31/22 | 54 | 364 | 364 | 0 | 1 | 0 | 0 | 3 |
| [57] † Eudimorphodon ranzii Last Modified in 10/31/22 | 229 | 190 | 190 | 0 | 0 | 0 | 0 | 3 |
| [58] † Dimorphodon macronyx Last Modified in 10/31/22 | 219 | 200 | 200 | 0 | 0 | 0 | 0 | 3 |
| [59] † Lagerpeton chanarensis Last Modified in 10/31/22 | 288 | 131 | 131 | 0 | 0 | 0 | 0 | 3 |
| [60] † Dromomeron gregorii Last Modified in 10/31/22 | 376 | 43 | 43 | 0 | 0 | 0 | 0 | 3 |
| [61] † Dromomeron romeri Last Modified in 10/31/22 | 356 | 63 | 63 | 0 | 0 | 0 | 0 | 3 |
| [62] † Marasuchus lilloensis Last Modified in 10/31/22 | 215 | 204 | 204 | 0 | 0 | 0 | 0 | 3 |
| [63] † Eucoelophysis baldwini Last Modified in 10/31/22 | 392 | 27 | 27 | 0 | 0 | 0 | 0 | 3 |
| [64] † Sacisaurus agudoensis Last Modified in 10/31/22 | 327 | 92 | 92 | 0 | 0 | 0 | 0 | 3 |
| [65] † Lewisuchus admixtus Last Modified in 10/31/22 | 286 | 133 | 133 | 0 | 0 | 0 | 0 | 3 |
| [66] † Pseudolagosuchus majori Last Modified in 10/31/22 | 322 | 97 | 97 | 0 | 0 | 0 | 0 | 3 |
| [67] † Lewisuchus pseudolagosuchus Last Modified in 10/31/22 | 191 | 228 | 228 | 0 | 0 | 0 | 0 | 3 |
| [68] † Eocursor parvus Last Modified in 10/31/22 | 276 | 142 | 142 | 0 | 1 | 0 | 0 | 3 |
| [69] † Silesaurus opolensis Last Modified in 10/31/22 | 80 | 340 | 339 | 0 | 0 | 0 | 0 | 3 |
| [70] † Pisanosaurus mertii Last Modified in 10/31/22 | 331 | 88 | 88 | 0 | 0 | 0 | 0 | 3 |
| [71] † Heterodontosaurus tucki Last Modified in 10/31/22 | 74 | 346 | 344 | 0 | 1 | 0 | 0 | 3 |
| [72] † Lesothosaurus diagnosticus Last Modified in 10/31/22 | 88 | 331 | 330 | 0 | 1 | 0 | 0 | 3 |
| [73] † Scutellosaurus lawleri Last Modified in 10/31/22 | 248 | 171 | 170 | 0 | 1 | 0 | 0 | 3 |
| [74] † Herrerasaurus ischigualastensis Last Modified in 10/31/22 | 69 | 349 | 349 | 0 | 1 | 0 | 0 | 3 |
| [75] † Staurikosaurus pricei Last Modified in 10/31/22 | 301 | 117 | 117 | 0 | 1 | 0 | 0 | 3 |
| [76] † Eoraptor lunensis Last Modified in 10/31/22 | 134 | 285 | 284 | 0 | 1 | 0 | 0 | 3 |
| [77] † Saturnalia tupiniquim Last Modified in 10/31/22 | 222 | 196 | 196 | 0 | 1 | 0 | 0 | 3 |
| [78] † Plateosaurus engelhardti Last Modified in 10/31/22 | 30 | 388 | 388 | 0 | 1 | 0 | 0 | 3 |
| [79] † Efraasia minor Last Modified in 10/31/22 | 95 | 323 | 323 | 0 | 1 | 0 | 0 | 3 |
| [80] † Tawa hallae Last Modified in 10/31/22 | 71 | 347 | 347 | 0 | 1 | 0 | 0 | 3 |
| [81] † Coelophysis bauri Last Modified in 10/31/22 | 52 | 366 | 366 | 0 | 1 | 0 | 0 | 3 |
| [82] † Dilophosaurus wetherelli Last Modified in 10/31/22 | 76 | 342 | 342 | 0 | 1 | 0 | 0 | 3 |
| [83] † Allosaurus fragilis Last Modified in 10/31/22 | 41 | 377 | 377 | 0 | 1 | 0 | 0 | 3 |
| [84] † Velociraptor mongoliensis Last Modified in 10/31/22 | 83 | 336 | 335 | 0 | 1 | 0 | 0 | 3 |
| [85] † Parringtonia gracilis Last Modified in 10/31/22 | 381 | 38 | 38 | 0 | 0 | 0 | 0 | 3 |
| [86] † Erpetosuchus granti Last Modified in 10/31/22 | 266 | 153 | 153 | 0 | 0 | 0 | 0 | 3 |
| [87] † Asilisaurus kongwe Last Modified in 10/31/22 | 227 | 177 | 177 | 0 | 15 | 0 | 0 | 3 |
| [88] † Asilisaurus kongwe skeleton Last Modified in 10/31/22 | 165 | 238 | 238 | 0 | 16 | 0 | 0 | 3 |
| [89] † Asilisaurus kongwe combined Last Modified in 10/31/22 | 136 | 268 | 267 | 0 | 16 | 0 | 0 | 3 |
| [90] † Teleocrater holotype Last Modified in 10/31/22 | 338 | 81 | 81 | 0 | 0 | 0 | 0 | 3 |
| [91] † Teleocrater 2015 quarry Last Modified in 10/31/22 | 258 | 161 | 161 | 0 | 0 | 0 | 0 | 3 |
| [92] † Teleocrater combined Last Modified in 10/31/22 | 248 | 171 | 171 | 0 | 0 | 0 | 0 | 3 |
| [93] † Dongusuchus efremovi Last Modified in 10/31/22 | 391 | 28 | 28 | 0 | 0 | 0 | 0 | 3 |
| [94] † Spondylosoma absconditum Last Modified in 10/31/22 | 382 | 37 | 37 | 0 | 0 | 0 | 0 | 3 |
| [95] † Yarasuchus deccanensis Last Modified in 10/31/22 | 287 | 129 | 129 | 0 | 3 | 0 | 0 | 3 |
| [96] † Scleromochlus taylori Last Modified in 10/31/22 | 334 | 85 | 85 | 0 | 0 | 0 | 0 | 3 |
| [97] † Mandasuchus total Last Modified in 10/31/22 | 262 | 155 | 154 | 0 | 3 | 0 | 0 | 3 |
| [98] † Nundasuchus Last Modified in 10/31/22 | 224 | 191 | 191 | 0 | 4 | 0 | 0 | 3 |
| [99] † Trialestes romeri Last Modified in 10/31/22 | 249 | 168 | 166 | 0 | 4 | 0 | 0 | 3 |
Project views
| type | number of views | Individual items viewed (where applicable) |
| Total project views | 6064 | |
| Matrix views | 257 | Matrix landing page (232 views); Matrix from Yanez et al 2021 - (prepared by MorphoBank curators from original submission) (25 views); |
| Project overview | 388 | |
| Media views | 950 | Media search (801 views); M848027 (149 views); |
| Documents list | 224 | |
| Bibliography | 206 | |
| Views for media list | 295 | |
| Specimen list | 460 | |
| Taxon list | 3284 |
Project downloads
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 4 | |
| Matrix downloads | 3 | Matrix from Yanez et al 2021 - (prepared by MorphoBank curators from original submission) (3 downloads); |
| Project downloads | 1 |