Project 2626: J. N. Choiniere, C. A. Forster, W. J. de Klerk. 2012. New information on Nqwebasaurus thwazi, a coelurosaurian theropod from the Early Cretaceous Kirkwood Formation in South Africa. Journal of African Earth Sciences. 71:1-17.
Abstract
We performed additional preparation on the holotype skeleton of Nqwebasaurus thwazi and discovered new skeletal material. We describe this material, which includes a maxilla with small, conical, unserrated teeth and bones of the braincase, as well as parts of the holotype postcranial anatomy that were previously poorly documented. We incorporate this new anatomical information into a broadly sampled matrix designed to test theropod relationships. Our phylogenetic results hypothesize that Nqwebasaurus is the basalmost ornithomimosaur, and recover numerous characters supporting this relationship, including features of the maxilla, frontal, dentition, axial skeleton, forelimb and hindlimb. Nqwebasaurus is the first African ornithomimosaur and the first Gondwanan member of this group known from articulated skeletal material, supporting the hypothesis that coelurosaurian groups were cosmopolitan during their early evolutionary history. The presence of reduced dentition and a gastric mill in Nqwebasaurus strongly suggest that this taxon was herbivorous.Read the article »
Article DOI: 10.1016/j.jafrearsci.2012.05.005
Project DOI: 10.7934/P2626, http://dx.doi.org/10.7934/P2626
| This project contains | Matrices |
|---|---|
Download Project SDD File  | Total scored cells: 25550 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 568 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 1290 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:510/58 |
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MorphoBank Project 2626
MorphoBank Project 2626
- Creation Date:
23 February 2017 - Publication Date:
23 February 2017 - Project views: 39882
- Media downloads: 2
- Matrix downloads: 10
Authors' Institutions
- Albany Museum, Grahamstown
- The George Washington University
- University of the Witwatersrand
- American Museum of Natural History
- Rhodes University
Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| MorphoBank Curator Project Administrator | 97 | 11 | 10 | 568 | 0 | 0 | 25550 (24118, 0, 1432) | 0 | 0 | 0 | ||||
| Jonah Choiniere Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| Catherine Forster Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M24524' (97 Taxa)
| taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] † Herrerasaurus ischigualastensis Last Modified in 02/23/17 | 189 | 353 | 352 | 0 | 26 | 0 | 0 | 3 |
| [2] † Nqwebasaurus thwazi Last Modified in 02/23/17 | 325 | 229 | 226 | 0 | 14 | 0 | 0 | 3 |
| [3] † Acrocanthosaurus atokensis Last Modified in 02/23/17 | 258 | 303 | 300 | 0 | 8 | 0 | 0 | 3 |
| [4] † Afrovenator abakensis Last Modified in 02/23/17 | 315 | 247 | 246 | 0 | 7 | 0 | 0 | 3 |
| [5] † Albertosaurus sarcophagus Last Modified in 02/23/17 | 160 | 401 | 401 | 0 | 7 | 0 | 0 | 3 |
| [6] † Allosaurus fragilis Last Modified in 02/23/17 | 45 | 510 | 501 | 0 | 15 | 0 | 0 | 3 |
| [7] † Alvarezsaurus calvoi Last Modified in 02/23/17 | 448 | 111 | 101 | 0 | 9 | 0 | 0 | 3 |
| [8] † Alxasaurus elesitaiensis Last Modified in 02/23/17 | 386 | 178 | 177 | 0 | 4 | 0 | 0 | 3 |
| [9] † Anserimimus planinychus Last Modified in 02/23/17 | 460 | 103 | 103 | 0 | 5 | 0 | 0 | 3 |
| [10] † Apsaravis ukhaana Last Modified in 02/23/17 | 345 | 197 | 197 | 0 | 26 | 0 | 0 | 3 |
| [11] † Archaeopteryx lithographica Last Modified in 02/23/17 | 197 | 349 | 348 | 0 | 22 | 0 | 0 | 3 |
| [12] † Avimimus portentosus Last Modified in 02/23/17 | 254 | 301 | 301 | 0 | 13 | 0 | 0 | 3 |
| [13] † Beipiaosaurus inexpectus Last Modified in 02/23/17 | 443 | 116 | 116 | 0 | 9 | 0 | 0 | 3 |
| [14] † Beishanlong grandis Last Modified in 02/23/17 | 468 | 100 | 100 | 0 | 0 | 0 | 0 | 3 |
| [15] † Buitreraptor gonzalezorum Last Modified in 02/23/17 | 383 | 179 | 179 | 0 | 6 | 0 | 0 | 3 |
| [16] † Byronosaurus jaffeei Last Modified in 02/23/17 | 395 | 160 | 159 | 0 | 13 | 0 | 0 | 3 |
| [17] † Carnotaurus sastrei Last Modified in 02/23/17 | 221 | 326 | 325 | 0 | 21 | 0 | 0 | 3 |
| [18] † Caudipteryx zhoui Last Modified in 02/23/17 | 304 | 252 | 247 | 0 | 13 | 0 | 0 | 3 |
| [19] † Ceratosaurus nasicornis Last Modified in 02/23/17 | 168 | 386 | 385 | 0 | 14 | 0 | 0 | 3 |
| [20] † Chirostenotes pergracilis Last Modified in 02/23/17 | 380 | 175 | 175 | 0 | 13 | 0 | 0 | 3 |
| [21] † Citipati osmolskae Last Modified in 02/23/17 | 182 | 337 | 336 | 0 | 49 | 0 | 0 | 3 |
| [22] † Coelophysis bauri Last Modified in 02/23/17 | 227 | 323 | 321 | 0 | 19 | 0 | 0 | 3 |
| [23] † Compsognathus longipes Last Modified in 02/23/17 | 230 | 322 | 316 | 0 | 16 | 0 | 0 | 3 |
| [24] † Conchoraptor gracilis Last Modified in 02/23/17 | 372 | 179 | 179 | 0 | 17 | 0 | 0 | 3 |
| [25] † Confuciusornis sanctus Last Modified in 02/23/17 | 273 | 246 | 246 | 0 | 49 | 0 | 0 | 3 |
| [26] † Cryolophosaurus ellioti Last Modified in 02/23/17 | 414 | 150 | 147 | 0 | 6 | 0 | 0 | 3 |
| [27] † Daspletosaurus torosus Last Modified in 02/23/17 | 358 | 202 | 201 | 0 | 8 | 0 | 0 | 3 |
| [28] † Deinonychus antirrhopus Last Modified in 02/23/17 | 134 | 420 | 420 | 0 | 14 | 0 | 0 | 3 |
| [29] † Dilong paradoxus Last Modified in 02/23/17 | 228 | 324 | 321 | 0 | 16 | 0 | 0 | 3 |
| [30] † Dilophosaurus wetherilli Last Modified in 02/23/17 | 206 | 348 | 345 | 0 | 16 | 0 | 0 | 3 |
| [31] † Dromaeosaurus albertensis Last Modified in 02/23/17 | 399 | 159 | 159 | 0 | 10 | 0 | 0 | 3 |
| [32] † EK troodontid Last Modified in 02/23/17 | 516 | 50 | 50 | 0 | 2 | 0 | 0 | 3 |
| [33] † Eoraptor lunensis Last Modified in 02/23/17 | 323 | 222 | 222 | 0 | 23 | 0 | 0 | 3 |
| [34] † Epidexipteryx hui Last Modified in 02/23/17 | 389 | 171 | 171 | 0 | 8 | 0 | 0 | 3 |
| [35] † Erlikosaurus andrewsi Last Modified in 02/23/17 | 367 | 179 | 178 | 0 | 22 | 0 | 0 | 3 |
| [36] † Eustreptospondylus oxiensis Last Modified in 02/23/17 | 195 | 357 | 355 | 0 | 17 | 0 | 0 | 3 |
| [37] † Falcarius utahensis Last Modified in 02/23/17 | 185 | 378 | 376 | 0 | 6 | 0 | 0 | 3 |
| [38] † Gallimimus bullatus Last Modified in 02/23/17 | 192 | 338 | 338 | 0 | 38 | 0 | 0 | 3 |
| [39] † Garudimimus brevipes Last Modified in 02/23/17 | 196 | 326 | 326 | 0 | 46 | 0 | 0 | 3 |
| [40] † Giganotosaurus carolinii Last Modified in 02/23/17 | 373 | 193 | 191 | 0 | 4 | 0 | 0 | 3 |
| [41] † Gigantoraptor erlianensis Last Modified in 02/23/17 | 467 | 86 | 86 | 0 | 15 | 0 | 0 | 3 |
| [42] † Gorgosaurus libratus Last Modified in 02/23/17 | 308 | 255 | 255 | 0 | 5 | 0 | 0 | 3 |
| [43] † Guanlong wucaii Last Modified in 02/23/17 | 68 | 487 | 476 | 0 | 14 | 0 | 0 | 3 |
| [44] † Haplocheirus sollers Last Modified in 02/23/17 | 109 | 444 | 437 | 0 | 16 | 0 | 0 | 3 |
| [45] † Harpymimus okladnikovi Last Modified in 02/23/17 | 226 | 306 | 305 | 0 | 36 | 0 | 0 | 3 |
| [46] † Huaxiagnathus orientalis Last Modified in 02/23/17 | 313 | 242 | 242 | 0 | 13 | 0 | 0 | 3 |
| [47] † Incisivosaurus gauthieri Last Modified in 02/23/17 | 348 | 207 | 207 | 0 | 13 | 0 | 0 | 3 |
| [48] † Limusaurus inextricabilis Last Modified in 02/23/17 | 213 | 301 | 298 | 0 | 56 | 0 | 0 | 3 |
| [49] † Linheraptor exquisitus Last Modified in 02/23/17 | 321 | 236 | 236 | 0 | 11 | 0 | 0 | 3 |
| [50] † Mahakala omnogovae Last Modified in 02/23/17 | 478 | 84 | 83 | 0 | 6 | 0 | 0 | 3 |
| [51] † Majungasaurus crenatissimus Last Modified in 02/23/17 | 164 | 392 | 391 | 0 | 12 | 0 | 0 | 3 |
| [52] † Mapusaurus roseae Last Modified in 02/23/17 | 380 | 186 | 186 | 0 | 2 | 0 | 0 | 3 |
| [53] † Masiakasaurus knopfleri Last Modified in 02/23/17 | 385 | 172 | 167 | 0 | 11 | 0 | 0 | 3 |
| [54] † Megalosaurus bucklandii Last Modified in 02/23/17 | 376 | 185 | 184 | 0 | 8 | 0 | 0 | 3 |
| [55] † Mei long Last Modified in 02/23/17 | 242 | 301 | 301 | 0 | 25 | 0 | 0 | 3 |
| [56] † Microraptor gui Last Modified in 02/23/17 | 330 | 223 | 223 | 0 | 15 | 0 | 0 | 3 |
| [57] † Microvenator celer Last Modified in 02/23/17 | 382 | 179 | 176 | 0 | 8 | 0 | 0 | 3 |
| [58] † Monolophosaurus jiangi Last Modified in 02/23/17 | 265 | 291 | 290 | 0 | 12 | 0 | 0 | 3 |
| [59] † Mononykus olecranus Last Modified in 02/23/17 | 319 | 228 | 228 | 0 | 21 | 0 | 0 | 3 |
| [60] † Neovenator salerii Last Modified in 02/23/17 | 261 | 303 | 301 | 0 | 4 | 0 | 0 | 3 |
| [61] † Nothronychus graffami Last Modified in 02/23/17 | 384 | 180 | 180 | 0 | 4 | 0 | 0 | 3 |
| [62] † Ornitholestes hermanni Last Modified in 02/23/17 | 224 | 330 | 325 | 0 | 15 | 0 | 0 | 3 |
| [63] † Ornithomimus edmontonicus Last Modified in 02/23/17 | 264 | 283 | 283 | 0 | 21 | 0 | 0 | 3 |
| [64] † Oviraptor philoceratops Last Modified in 02/23/17 | 433 | 120 | 120 | 0 | 15 | 0 | 0 | 3 |
| [65] † Parvicursor remotus Last Modified in 02/23/17 | 439 | 112 | 109 | 0 | 19 | 0 | 0 | 3 |
| [66] † Patagonykus puertai Last Modified in 02/23/17 | 408 | 155 | 142 | 0 | 6 | 0 | 0 | 3 |
| [67] † Pelecanimimus polyodon Last Modified in 02/23/17 | 317 | 237 | 230 | 0 | 14 | 0 | 0 | 3 |
| [68] † Piatnitzkysaurus floresi Last Modified in 02/23/17 | 395 | 169 | 168 | 0 | 4 | 0 | 0 | 3 |
| [69] † Proceratosaurus bradleyi Last Modified in 02/23/17 | 415 | 147 | 145 | 0 | 6 | 0 | 0 | 3 |
| [70] † Rahonavis ostromi Last Modified in 02/23/17 | 350 | 212 | 212 | 0 | 6 | 0 | 0 | 3 |
| [71] † Rinchenia mongoliensis Last Modified in 02/23/17 | 453 | 101 | 101 | 0 | 14 | 0 | 0 | 3 |
| [72] † Sapeornis chaoyangensis Last Modified in 02/23/17 | 259 | 278 | 278 | 0 | 31 | 0 | 0 | 3 |
| [73] † Saurornithoides mongoliensis Last Modified in 02/23/17 | 406 | 155 | 155 | 0 | 7 | 0 | 0 | 3 |
| [74] † Saurornitholestes langstoni Last Modified in 02/23/17 | 363 | 201 | 196 | 0 | 4 | 0 | 0 | 3 |
| [75] † Segnosaurus galboensis Last Modified in 02/23/17 | 424 | 141 | 141 | 0 | 3 | 0 | 0 | 3 |
| [76] † Shenzhousaurus orientalis Last Modified in 02/23/17 | 313 | 225 | 225 | 0 | 30 | 0 | 0 | 3 |
| [77] † Shuvuuia deserti Last Modified in 02/23/17 | 78 | 442 | 438 | 0 | 48 | 0 | 0 | 3 |
| [78] † Sinornithoides youngi Last Modified in 02/23/17 | 403 | 157 | 157 | 0 | 8 | 0 | 0 | 3 |
| [79] † Sinornithosaurus millenii Last Modified in 02/23/17 | 323 | 232 | 232 | 0 | 13 | 0 | 0 | 3 |
| [80] † Sinosauropteryx prima Last Modified in 02/23/17 | 262 | 292 | 292 | 0 | 14 | 0 | 0 | 3 |
| [81] † Sinovenator changii Last Modified in 02/23/17 | 316 | 248 | 248 | 0 | 4 | 0 | 0 | 3 |
| [82] † Sinraptor dongi Last Modified in 02/23/17 | 109 | 445 | 441 | 0 | 14 | 0 | 0 | 3 |
| [83] † Stokesosaurus langhami Last Modified in 02/23/17 | 445 | 122 | 118 | 0 | 1 | 0 | 0 | 3 |
| [84] † Struthiomimus altus Last Modified in 02/23/17 | 146 | 376 | 376 | 0 | 46 | 0 | 0 | 3 |
| [85] † Suchomimus tenerensis Last Modified in 02/23/17 | 375 | 187 | 185 | 0 | 8 | 0 | 0 | 3 |
| [86] † Syntarsus rhodesiensis Last Modified in 02/23/17 | 180 | 371 | 368 | 0 | 18 | 0 | 0 | 3 |
| [87] † Tanycolagreus topwilsoni Last Modified in 02/23/17 | 328 | 237 | 235 | 0 | 3 | 0 | 0 | 3 |
| [88] † Tarbosaurus bataar Last Modified in 02/23/17 | 361 | 199 | 199 | 0 | 8 | 0 | 0 | 3 |
| [89] † Tawa hallae Last Modified in 02/23/17 | 284 | 264 | 264 | 0 | 20 | 0 | 0 | 3 |
| [90] † Torvosaurus tanneri Last Modified in 02/23/17 | 344 | 215 | 214 | 0 | 9 | 0 | 0 | 3 |
| [91] † Troodon formosus Last Modified in 02/23/17 | 269 | 284 | 281 | 0 | 15 | 0 | 0 | 3 |
| [92] † Tsaagan mangas Last Modified in 02/23/17 | 350 | 205 | 205 | 0 | 13 | 0 | 0 | 3 |
| [93] † Tyrannosaurus rex Last Modified in 02/23/17 | 32 | 518 | 516 | 0 | 18 | 0 | 0 | 3 |
| [94] † Unenlagia comahuensis Last Modified in 02/23/17 | 446 | 120 | 112 | 0 | 2 | 0 | 0 | 3 |
| [95] † Velociraptor mongoliensis Last Modified in 02/23/17 | 118 | 431 | 431 | 0 | 19 | 0 | 0 | 3 |
| [96] † Zanabazar junior Last Modified in 02/23/17 | 378 | 179 | 179 | 0 | 11 | 0 | 0 | 3 |
| [97] † Zuolong salleei Last Modified in 02/23/17 | 321 | 240 | 237 | 0 | 7 | 0 | 0 | 3 |
Project views
| type | number of views | Individual items viewed (where applicable) |
| Total project views | 39882 | |
| Project overview | 1895 | |
| Media views | 10383 | Media search (3255 views); M437343 (711 views); M437346 (760 views); M437344 (939 views); M437348 (610 views); M437340 (694 views); M437342 (689 views); M437345 (636 views); M437347 (652 views); M437349 (704 views); M437341 (733 views); |
| Matrix views | 1558 | Matrix landing page (1380 views); matrix (178 views); |
| Taxon list | 13133 | |
| Documents list | 1032 | |
| Bibliography | 837 | |
| Specimen list | 9082 | |
| Views for media list | 1960 | |
| Folio views | 2 | Folio list (2 views); |
Project downloads
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 377 | |
| Project downloads | 358 | |
| Document downloads | 7 | Matrix (3 downloads); character names and states (4 downloads); |
| Matrix downloads | 10 | matrix (10 downloads); |
| Media downloads | 2 | M437349 (2 downloads); |