Project 987: V. Vahtera, G. Edgecombe, G. Giribet. 2013. Phylogenetics of scolopendromorph centipedes: Can denser taxon sampling improve an artificial classification?. Invertebrate Systematics. 27 (5):578-602.
Abstract
Previous phylogenetic analyses of the centipede order Scolopendromorpha indicated a fundamental division into blind and ocellate clades. These analyses corroborated the monophyly of most families and tribes but suggested that several species-rich, cosmopolitan genera in traditional and current classifications are polyphyletic. Denser taxon sampling is applied to a dataset of 122 morphological characters and sequences for four nuclear and mitochondrial loci. Phylogenetic analyses including 98 species and subspecies of Scolopendromorpha employ parsimony under dynamic and static homology schemes as well as maximum likelihood and Bayesian inference of multiple sequence alignments. The monotypic Australian genera Notiasemus and Kanparka nest within Cormocephalus and Scolopendra, respectively, and the New Caledonian Campylostigmus is likewise a clade within Cormocephalus. New World Scolopendra are more closely related to Hemiscolopendra and Arthrorhabdus than to Scolopendra s.s., which is instead closely allied to Asanada; the tribe Asanadini nests within Scolopendrini for molecular and combined datasets. The generic classification of Otostigmini has a poor fit to phylogenetic relationships, although nodal support within this tribe is weak. New synonymies are proposed for Ectonocryptopinae Shelley & Mercurio, 2005 (= Newportiinae Pocock, 1896), Asanadini Verhoeff, 1907 (= Scolopendrini Leach, 1814), and Kanparka Waldock & Edgecombe, 2012 (= Scolopendra Linnaeus, 1758). Scolopendrid systematics largely depicts incongruence between phylogeny and classification rather than between morphology and molecules.Read the article »
Project DOI: 10.7934/P987, http://dx.doi.org/10.7934/P987
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 10464 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 122 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 277 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:122/0 |
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MorphoBank Project 987
MorphoBank Project 987
- Creation Date:
10 July 2013 - Publication Date:
05 November 2013 - Project views: 39579
- Matrix downloads: 27
Authors' Institutions
- Natural History Museum, London
- Harvard University
- University of Turku
- Finnish Museum of Natural History
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 108 | 1 | 1 | 122 | 0 | 0 | 10464 (9648, 0, 816) | 0 | 0 | 0 | ||||
Greg Edgecombe Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Gonzalo Giribet Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M2184' (108 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Scutigera coleoptrata Last Modified in 12/05/13 | 6 | 86 | 86 | 0 | 30 | 0 | 0 | 3 |
[2] Lithobius forficatus Last Modified in 12/05/13 | 4 | 89 | 89 | 0 | 29 | 0 | 0 | 3 |
[3] Craterostigmus tasmanianus Last Modified in 12/05/13 | 26 | 71 | 71 | 0 | 25 | 0 | 0 | 3 |
[4] Dicellophilus carniolensis Last Modified in 12/05/13 | 15 | 78 | 78 | 0 | 29 | 0 | 0 | 3 |
[5] Bothriogaster signata Last Modified in 12/05/13 | 35 | 69 | 69 | 0 | 18 | 0 | 0 | 3 |
[6] Plesioschendyla confossa Last Modified in 12/05/13 | 39 | 67 | 67 | 0 | 16 | 0 | 0 | 3 |
[7] Plutonium zwierleini Last Modified in 12/05/13 | 6 | 107 | 107 | 0 | 9 | 0 | 0 | 3 |
[8] Theatops erythrocephalus Last Modified in 12/05/13 | 2 | 111 | 111 | 0 | 9 | 0 | 0 | 3 |
[9] Theatops posticus Last Modified in 12/05/13 | 4 | 108 | 108 | 0 | 10 | 0 | 0 | 3 |
[10] C. (cryptops) anomalans Last Modified in 12/05/13 | 18 | 92 | 92 | 0 | 12 | 0 | 0 | 3 |
[11] C. (cryptops) australis Last Modified in 12/05/13 | 8 | 102 | 102 | 0 | 12 | 0 | 0 | 3 |
[12] C. (cryptops) doriae Last Modified in 12/05/13 | 37 | 74 | 74 | 0 | 11 | 0 | 0 | 3 |
[13] C. (cryptops) hortensis Last Modified in 12/05/13 | 2 | 106 | 106 | 0 | 14 | 0 | 0 | 3 |
[14] C. (cryptops) lamprethus Last Modified in 12/05/13 | 29 | 81 | 81 | 0 | 12 | 0 | 0 | 3 |
[15] C. (cryptops) niuhensis Last Modified in 12/05/13 | 43 | 71 | 71 | 0 | 8 | 0 | 0 | 3 |
[16] C. (cryptops) parisi Last Modified in 12/05/13 | 40 | 74 | 74 | 0 | 8 | 0 | 0 | 3 |
[17] C. (cryptops) punicus Last Modified in 12/05/13 | 28 | 82 | 82 | 0 | 12 | 0 | 0 | 3 |
[18] C. (cryptops) trisulcatus Last Modified in 12/05/13 | 32 | 79 | 79 | 0 | 11 | 0 | 0 | 3 |
[19] C. (cryptops) typhloporus Last Modified in 12/05/13 | 35 | 78 | 78 | 0 | 9 | 0 | 0 | 3 |
[20] C. (paracryptops) indicus Last Modified in 12/05/13 | 38 | 76 | 75 | 0 | 9 | 0 | 0 | 3 |
[21] C. (paracryptops) weberi Last Modified in 12/05/13 | 12 | 100 | 99 | 0 | 11 | 0 | 0 | 3 |
[22] C. (trigonocryptops) galatheae Last Modified in 12/05/13 | 45 | 69 | 69 | 0 | 8 | 0 | 0 | 3 |
[23] C. (trigonocryptops) pictus Last Modified in 12/05/13 | 35 | 78 | 78 | 0 | 9 | 0 | 0 | 3 |
[24] C. (trigonocryptops) sarasini Last Modified in 12/05/13 | 44 | 70 | 70 | 0 | 8 | 0 | 0 | 3 |
[25] C. (trigonocryptops) spinipes Last Modified in 12/05/13 | 11 | 101 | 101 | 0 | 10 | 0 | 0 | 3 |
[26] Kethops utahensis Last Modified in 12/05/13 | 12 | 102 | 102 | 0 | 8 | 0 | 0 | 3 |
[27] N. (ectonocryptoides) quadrimeropus Last Modified in 12/05/13 | 12 | 100 | 100 | 0 | 10 | 0 | 0 | 3 |
[28] Newportia adisi Last Modified in 12/05/13 | 47 | 69 | 69 | 0 | 6 | 0 | 0 | 3 |
[29] Newportia divergens Last Modified in 12/05/13 | 12 | 103 | 103 | 0 | 7 | 0 | 0 | 3 |
[30] Newportia ernsti ernsti Last Modified in 12/05/13 | 32 | 84 | 84 | 0 | 6 | 0 | 0 | 3 |
[31] Newportia longitarsis stechowi Last Modified in 12/05/13 | 12 | 103 | 102 | 0 | 8 | 0 | 0 | 3 |
[32] Newportia monticola Last Modified in 12/05/13 | 7 | 107 | 107 | 0 | 8 | 0 | 0 | 3 |
[33] Newportia oreina Last Modified in 12/05/13 | 47 | 71 | 71 | 0 | 4 | 0 | 0 | 3 |
[34] Newportia stollii Last Modified in 12/05/13 | 10 | 105 | 105 | 0 | 7 | 0 | 0 | 3 |
[35] N. (tidops) collaris Last Modified in 12/05/13 | 14 | 100 | 100 | 0 | 8 | 0 | 0 | 3 |
[36] Scolopocryptops macrodon Last Modified in 12/05/13 | 46 | 71 | 71 | 0 | 5 | 0 | 0 | 3 |
[37] Scolopocryptops melanostoma Last Modified in 12/05/13 | 28 | 87 | 87 | 0 | 7 | 0 | 0 | 3 |
[38] Scolopocryptops mexicana Last Modified in 12/05/13 | 2 | 112 | 112 | 0 | 8 | 0 | 0 | 3 |
[39] Scolopocryptops miersii Last Modified in 12/05/13 | 4 | 110 | 110 | 0 | 8 | 0 | 0 | 3 |
[40] Scolopocryptops nipponicus Last Modified in 12/05/13 | 47 | 70 | 70 | 0 | 5 | 0 | 0 | 3 |
[41] Scolopocryptops nigridius Last Modified in 12/05/13 | 35 | 81 | 81 | 0 | 6 | 0 | 0 | 3 |
[42] Scolopocryptops rubiginosus Last Modified in 12/05/13 | 24 | 91 | 91 | 0 | 7 | 0 | 0 | 3 |
[43] Scolopocryptops sexspinosus Last Modified in 12/05/13 | 43 | 74 | 74 | 0 | 5 | 0 | 0 | 3 |
[44] Scolopocryptops spinicauda Last Modified in 12/05/13 | 11 | 104 | 104 | 0 | 7 | 0 | 0 | 3 |
[45] Scolopendra afer Last Modified in 12/05/13 | 38 | 81 | 81 | 0 | 3 | 0 | 0 | 3 |
[46] Scolopendra canidens Last Modified in 12/05/13 | 3 | 110 | 110 | 0 | 9 | 0 | 0 | 3 |
[47] Scolopendra cingulata Last Modified in 12/05/13 | 3 | 110 | 110 | 0 | 9 | 0 | 0 | 3 |
[48] Scolopendra dehaani Last Modified in 12/05/13 | 34 | 84 | 84 | 0 | 4 | 0 | 0 | 3 |
[49] Scolopendra hardwickei Last Modified in 12/05/13 | 45 | 74 | 74 | 0 | 3 | 0 | 0 | 3 |
[50] Scolopendra laeta Last Modified in 12/05/13 | 4 | 109 | 109 | 0 | 9 | 0 | 0 | 3 |
[51] Scolopendra leki Last Modified in 12/05/13 | 35 | 84 | 84 | 0 | 3 | 0 | 0 | 3 |
[52] Scolopendra mirabilis Last Modified in 12/05/13 | 31 | 88 | 88 | 0 | 3 | 0 | 0 | 3 |
[53] Scolopendra morsitans Last Modified in 12/05/13 | 1 | 112 | 112 | 0 | 9 | 0 | 0 | 3 |
[54] Scolopendra multidens Last Modified in 12/05/13 | 36 | 83 | 83 | 0 | 3 | 0 | 0 | 3 |
[55] Scolopendra oraniensis Last Modified in 12/05/13 | 3 | 110 | 110 | 0 | 9 | 0 | 0 | 3 |
[56] Scolopendra subspinipes mutilans Last Modified in 12/05/13 | 41 | 78 | 78 | 0 | 3 | 0 | 0 | 3 |
[57] Scolopendra subspinipes subspinipes Last Modified in 12/05/13 | 24 | 95 | 95 | 0 | 3 | 0 | 0 | 3 |
[58] Nuretiniella alternans Last Modified in 12/05/13 | 30 | 89 | 89 | 0 | 3 | 0 | 0 | 3 |
[59] Nuretiniella polymorpha Last Modified in 12/05/13 | 36 | 83 | 83 | 0 | 3 | 0 | 0 | 3 |
[60] Nuretiniella valida Last Modified in 12/05/13 | 26 | 93 | 93 | 0 | 3 | 0 | 0 | 3 |
[61] Nuretiniella viridis Last Modified in 12/05/13 | 35 | 84 | 84 | 0 | 3 | 0 | 0 | 3 |
[62] Cormocephalus anceps Last Modified in 12/05/13 | 44 | 74 | 74 | 0 | 4 | 0 | 0 | 3 |
[63] Cormocephalus aurantiipes Last Modified in 12/05/13 | 3 | 109 | 109 | 0 | 10 | 0 | 0 | 3 |
[64] Cormocephalus bonaerius Last Modified in 12/05/13 | 35 | 82 | 82 | 0 | 5 | 0 | 0 | 3 |
[65] Cormocephalus calcaratus Last Modified in 12/05/13 | 33 | 85 | 85 | 0 | 4 | 0 | 0 | 3 |
[66] Cormocephalus elegans Last Modified in 12/05/13 | 45 | 73 | 73 | 0 | 4 | 0 | 0 | 3 |
[67] Cormocephalus gervaisianus Last Modified in 12/05/13 | 29 | 89 | 89 | 0 | 4 | 0 | 0 | 3 |
[68] Cormocephalus hartmeyeri Last Modified in 12/05/13 | 11 | 101 | 101 | 0 | 10 | 0 | 0 | 3 |
[69] Comocephalus longipes Last Modified in 12/05/13 | 33 | 85 | 85 | 0 | 4 | 0 | 0 | 3 |
[70] Cormocephalus monteithi Last Modified in 12/05/13 | 45 | 73 | 73 | 0 | 4 | 0 | 0 | 3 |
[71] Cormocephalus multispinus Last Modified in 12/05/13 | 33 | 85 | 85 | 0 | 4 | 0 | 0 | 3 |
[72] Cormocephalus neocaledonicus Last Modified in 12/05/13 | 38 | 80 | 80 | 0 | 4 | 0 | 0 | 3 |
[73] Cormocephalus nitidus Last Modified in 12/05/13 | 33 | 85 | 85 | 0 | 4 | 0 | 0 | 3 |
[74] Cormocephalus pseudopunctatus Last Modified in 12/05/13 | 36 | 82 | 82 | 0 | 4 | 0 | 0 | 3 |
[75] Cormocephalus westwoodi Last Modified in 12/05/13 | 32 | 86 | 86 | 0 | 4 | 0 | 0 | 3 |
[76] Campylostigmus decipiens Last Modified in 12/05/13 | 31 | 87 | 87 | 0 | 4 | 0 | 0 | 3 |
[77] Campylostigmus orientalis Last Modified in 12/05/13 | 26 | 92 | 92 | 0 | 4 | 0 | 0 | 3 |
[78] Akymnopellis chilensis Last Modified in 12/05/13 | 4 | 109 | 108 | 0 | 10 | 0 | 0 | 3 |
[79] Hemiscolopendra marginata Last Modified in 12/05/13 | 30 | 88 | 88 | 0 | 4 | 0 | 0 | 3 |
[80] Arthrorhabdus formosus Last Modified in 12/05/13 | 4 | 109 | 109 | 0 | 9 | 0 | 0 | 3 |
[81] Scolopendropsis bahiensis Last Modified in 12/05/13 | 12 | 101 | 100 | 0 | 10 | 0 | 0 | 3 |
[82] Notiasemus glauerti Last Modified in 12/05/13 | 12 | 99 | 99 | 0 | 11 | 0 | 0 | 3 |
[83] Asanada brevicornis Last Modified in 12/05/13 | 3 | 103 | 103 | 0 | 16 | 0 | 0 | 3 |
[84] Asanada socotrana Last Modified in 12/05/13 | 5 | 101 | 101 | 0 | 16 | 0 | 0 | 3 |
[85] Otostigmus angusticeps Last Modified in 12/05/13 | 33 | 86 | 86 | 0 | 3 | 0 | 0 | 3 |
[86] Otostigmus astenus Last Modified in 12/05/13 | 3 | 110 | 109 | 0 | 10 | 0 | 0 | 3 |
[87] Otostigmus multidens Last Modified in 12/05/13 | 33 | 86 | 86 | 0 | 3 | 0 | 0 | 3 |
[88] Otostigmus politus politus Last Modified in 12/05/13 | 34 | 85 | 85 | 0 | 3 | 0 | 0 | 3 |
[89] Otostigmus ruficeps Last Modified in 12/05/13 | 39 | 81 | 81 | 0 | 2 | 0 | 0 | 3 |
[90] Otostigmus rugulosus Last Modified in 12/05/13 | 26 | 94 | 93 | 0 | 3 | 0 | 0 | 3 |
[91] Otostigmus scaber Last Modified in 12/05/13 | 32 | 87 | 87 | 0 | 3 | 0 | 0 | 3 |
[92] Otostigmus spinicauda Last Modified in 12/05/13 | 25 | 95 | 94 | 0 | 3 | 0 | 0 | 3 |
[93] Parotostigmus caraibicus Last Modified in 12/05/13 | 31 | 86 | 86 | 0 | 5 | 0 | 0 | 3 |
[94] Digitipes barnabasi Last Modified in 12/05/13 | 39 | 81 | 81 | 0 | 2 | 0 | 0 | 3 |
[95] Digitipes coonoorensis Last Modified in 12/05/13 | 32 | 87 | 87 | 0 | 3 | 0 | 0 | 3 |
[96] Digitipes jonesii Last Modified in 12/05/13 | 32 | 87 | 87 | 0 | 3 | 0 | 0 | 3 |
[97] Alipes crotalus Last Modified in 12/05/13 | 4 | 106 | 106 | 0 | 12 | 0 | 0 | 3 |
[98] Alipes grandidieri Last Modified in 12/05/13 | 28 | 90 | 90 | 0 | 4 | 0 | 0 | 3 |
[99] Ethmostigmus curtipes Last Modified in 12/05/13 | 48 | 72 | 72 | 0 | 2 | 0 | 0 | 3 |
[100] Ethmostigmus rubripes rubripes Last Modified in 12/05/13 | 3 | 109 | 109 | 0 | 10 | 0 | 0 | 3 |
[101] Ethmostigmus trigonopodus Last Modified in 12/05/13 | 24 | 95 | 95 | 0 | 3 | 0 | 0 | 3 |
[102] Rhysida afra Last Modified in 12/05/13 | 35 | 85 | 84 | 0 | 3 | 0 | 0 | 3 |
[103] Rhysida cf carinulata Last Modified in 12/05/13 | 45 | 75 | 75 | 0 | 2 | 0 | 0 | 3 |
[104] Rhysida longicornis Last Modified in 12/05/13 | 46 | 74 | 74 | 0 | 2 | 0 | 0 | 3 |
[105] Rhysida nuda Last Modified in 12/05/13 | 1 | 111 | 111 | 0 | 10 | 0 | 0 | 3 |
[106] Rhysida polyacantha Last Modified in 12/05/13 | 33 | 86 | 86 | 0 | 3 | 0 | 0 | 3 |
[107] Edentistoma octosulcatus Last Modified in 12/05/13 | 10 | 98 | 98 | 0 | 14 | 0 | 0 | 3 |
[108] Sterropristes violaceus Last Modified in 12/05/13 | 44 | 74 | 74 | 0 | 4 | 0 | 0 | 3 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 39579 | |
Project overview | 5803 | |
Media views | 7001 | Media search (5984 views); M304087 (1017 views); |
Matrix views | 3087 | Matrix landing page (2794 views); Vahtera et al Scolopendromorph matrix (293 views); |
Specimen list | 3301 | |
Taxon list | 19125 | |
Bibliography | 1248 | |
Documents list | 14 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 423 | |
Matrix downloads | 27 | Vahtera et al Scolopendromorph matrix (27 downloads); |
Project downloads | 396 |