Project 683: Black, K. H., M. Archer, and S. J. Hand. 2012. New Tertiary Koala (Marsupialia, Phascolarctidae) from Riversleigh, Australia, with a Revision of Phascolarctid Phylogenetics, Paleoecology, and Paleobiodiversity. Journal of Vertebrate Paleontology. 32 (1):125-138.

Specimen: †Priscakoala lucyturnbulli (QM:F20303)
View: Partial left maxilla with M1-3, in occlusal view
View: Partial left maxilla with M1-3, in occlusal view
Abstract
Here we describe the most plesiomorphic koala yet known. Priscakoala lucyturnbullae, gen. et sp. nov., is the fourth and largest koala species described from Miocene deposits in the Riversleigh World Heritage area, northern Australia. It is known from a maxilla with M1–3 and isolated M2, M3 or M4 and m1. Relationships within the diprotodontian suborder Vombatiformes are clarified using cranial and dental characters based on a data set compiled from new and more complete cranial materials for each of the respective vombatiform families. Monophyly of Phascolarctidae is supported by the development of a protostylid and metastylid on m1. Priscakoala lucyturnbullae is the most plesiomorphic phascolarctid. Inclusion in the analysis of the enigmatic Pliocene genus Koobor, currently classified as Vombatiformes incertae sedis, indicates phascolarctomorphian affinities for the genus.Read the article »
| This project contains | Matrices |
|---|---|
| Total scored cells: 1013 Total media associated with cells: 0 Total labels associated with cell media: 0 |
| Characters | |
| Total characters: 71 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 180 Total character states with associated media: 0 Total character states with media with labels:0 |
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MorphoBank Project 683
MorphoBank Project 683
- Creation Date:
07 March 2012 - Publication Date:
01 April 2012 - Project views: 2871

- Project downloads: 61

Members
| member name | taxa |
specimens |
media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
| Natalie Milbrodt Project Administrator Last logged in 03/30/12 | 19 | 2 | 2 | 71 | 0 | 0 | 1013 (991, 0, 22) | 0 | 0 | 0 | ||||
| Maureen Admin Full membership Last logged in 03/08/12 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
| Karen Black Full membership Last logged in 03/30/12 | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Taxonomic Overview for Matrix 'M1586' (19 Taxa)
| taxon | unscored cells |
scored cells |
cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
| [1] †Galadi speciosus Last Modified in 03/30/12 | 3 | 67 | 0 | 0 | 1 | 0 | 0 | 3 |
| [2] Cercartetus lepidus Last Modified in 03/07/12 | 1 | 69 | 0 | 0 | 1 | 0 | 0 | 3 |
| [3] Phascolarctos cinereus Last Modified in 03/07/12 | 0 | 72 | 0 | 0 | 1 | 0 | 0 | 3 |
| [4] †Litokoala kutjamarpensis Last Modified in 03/30/12 | 14 | 56 | 0 | 0 | 1 | 0 | 0 | 3 |
| [5] †Nimiokoala greystanesi Last Modified in 03/30/12 | 11 | 59 | 0 | 0 | 1 | 0 | 0 | 3 |
| [6] †Perikoala robusta Last Modified in 03/30/12 | 32 | 38 | 0 | 0 | 1 | 0 | 0 | 3 |
| [7] †Madakoala devisi Last Modified in 03/30/12 | 28 | 42 | 0 | 0 | 1 | 0 | 0 | 3 |
| [8] †Invictokoala monticola Last Modified in 03/30/12 | 58 | 12 | 0 | 0 | 1 | 0 | 0 | 3 |
| [9] †Priscakoala lucyturnbulli Last Modified in 03/30/12 | 47 | 23 | 0 | 0 | 1 | 0 | 0 | 3 |
| [10] †Koobor notabilis Last Modified in 03/30/12 | 48 | 22 | 0 | 0 | 1 | 0 | 0 | 3 |
| [11] †Priscileo roskellyae Last Modified in 03/30/12 | 3 | 66 | 0 | 0 | 2 | 0 | 0 | 3 |
| [12] Vombatus ursinus Last Modified in 03/07/12 | 0 | 69 | 0 | 0 | 2 | 0 | 0 | 3 |
| [13] †Warendja wakefieldi Last Modified in 03/30/12 | 13 | 56 | 0 | 0 | 2 | 0 | 0 | 3 |
| [14] †Kuterintja ngama Last Modified in 03/30/12 | 30 | 40 | 0 | 0 | 1 | 0 | 0 | 3 |
| [15] †Namilamadeta albivenator Last Modified in 03/30/12 | 0 | 70 | 0 | 0 | 1 | 0 | 0 | 3 |
| [16] †Marada arcanum Last Modified in 03/30/12 | 45 | 25 | 0 | 0 | 1 | 0 | 0 | 3 |
| [17] †Propalorchestes novaculacephalus Last Modified in 03/30/12 | 7 | 63 | 0 | 0 | 1 | 0 | 0 | 3 |
| [18] †Ngapakaldia tedfordi Last Modified in 03/30/12 | 0 | 70 | 0 | 0 | 1 | 0 | 0 | 3 |
| [19] †Nimbadon lavarackorum Last Modified in 03/30/12 | 0 | 72 | 0 | 0 | 1 | 0 | 0 | 3 |
Project views 
| type | number of views | Individual items viewed (where applicable) |
| Total project views | 2871 | |
| Project overview | 343 | |
| Matrix views | 172 | Matrix landing page (156 views); Koala matrix (16 views); |
| Media views | 623 | Media search (458 views); M102645 (91 views); M102643 (74 views); |
| Views for media list | 405 | |
| Specimen list | 306 | |
| Taxon list | 737 | |
| Bibliography | 141 | |
| Documents list | 144 |
Project downloads 
| type | number of downloads | Individual items downloaded (where applicable) |
| Total downloads from project | 61 | |
| Media downloads | 35 | M102645 (23 downloads); M102643 (12 downloads); |
| Matrix downloads | 2 | Koala matrix (2 downloads); |
| Document downloads | 5 | Copyright information for exemplar media (5 downloads); |
| Project downloads | 19 |
