Project 610: I. M. Vea, D. A. Grimaldi. 2012. Phylogeny of ensign scale insects (Hemiptera: Coccoidea: Ortheziidae) based on the morphology of Recent and fossil females. Systematic Entomology. 37 (4):758-783.
Abstract
The Ortheziidae (ensign scale insects) is a morphologically well-definedfamily. The morphology and occurrence in the fossil record suggests a probable early origin of the family in scale insect evolution. The present phylogenetic analysis – based on 69 morphological characters of female ortheziids, using 39 exemplar Recent species – provides the first analytical assessment of relationships among living genera of the family, as well as the relationships of eight fossil species, based on complete, well-preserved specimens in amber. Monophyly of the subfamilies
Newsteadiinae, Ortheziinae and Ortheziolinae is supported, but Nipponortheziinae is found to be paraphyletic by inclusion of the Ortheziolinae. Thus, the subfamily Ortheziolinae is reduced in rank to tribe Ortheziolini stat.n., which now includes Matileortheziola Kozár & Foldi, Ortheziolacoccus Kozár, Ortheziolamameti Kozár and Ortheziola Sulc. Consequently, the tribes Matileortheziolini, Ortheziolacoccini and Ortheziolamametini are synonymized (syn.n.) here under Ortheziolini. Five new species and one new genus of fossil ensign scales are described from three amber deposits: Burmorthezia gen.n. with type species Burmorthezia kotejai sp.n. and also B. insolita sp.n., both in mid-Cretaceous Burmese amber (98 Ma) and Arctorthezia baltica sp.n. in Eocene Baltic amber (c. 43 Ma) based on second-instar nymphs; Mixorthezia kozari sp.n. and M. dominicana sp.n. in Miocene Dominican
amber (c. 17 Ma) based on adult females. Fossil placements are unambiguous, with Burmorthezia forming a stem to crown-group (Recent and Tertiary) Ortheziidae.
A summary of described fossil ortheziids is provided.
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Project DOI: 10.7934/P610, http://dx.doi.org/10.7934/P610
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 2969 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 69 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 172 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:69/0 |
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MorphoBank Project 610
MorphoBank Project 610
- Creation Date:
14 October 2011 - Publication Date:
13 September 2012 - Project views: 33344
- Media downloads: 1
- Matrix downloads: 27
Authors' Institutions
- American Museum of Natural History
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||||||||||
Isabelle Vea Project Administrator | 52 | 6 | 1 | 69 | 0 | 69 | 0 | 0 | 2969 (2920, 0, 49) | 0 | 0 | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M1475' (47 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Matsucoccus gallicolus Last Modified in 12/05/13 | 2 | 58 | 58 | 0 | 9 | 0 | 0 | 1 |
[2] Pseudococcus longispinus Last Modified in 12/05/13 | 0 | 61 | 61 | 0 | 8 | 0 | 0 | 1 |
[3] Icerya purchasi Last Modified in 12/05/13 | 0 | 62 | 62 | 0 | 7 | 0 | 0 | 1 |
[4] Puto yuccae Last Modified in 12/05/13 | 0 | 63 | 63 | 0 | 6 | 0 | 0 | 1 |
[5] Newsteadia floccosa Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[6] Newsteadia morrisoni Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[7] Newsteadia monikae Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[8] Newsteadia caledoniensis Last Modified in 12/05/13 | 1 | 68 | 68 | 0 | 0 | 0 | 0 | 1 |
[9] Newsteadia minima Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[10] Newsteadia americana Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[11] Newsteadia africana Last Modified in 12/05/13 | 1 | 68 | 68 | 0 | 0 | 0 | 0 | 1 |
[12] Jermycoccus boliviensis Last Modified in 12/05/13 | 0 | 67 | 67 | 0 | 2 | 0 | 0 | 1 |
[13] Neomixorthezia brazilana Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[14] Neomixorthezia machupicchui Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[15] Neonipponorthezia regina Last Modified in 12/05/13 | 9 | 60 | 60 | 0 | 0 | 0 | 0 | 1 |
[16] Nipponorthezia ardisiae Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[17] Nipponorthezia obscura Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[18] † Nipponorthezinella Last Modified in 12/05/13 | 30 | 39 | 39 | 0 | 0 | 0 | 0 | 1 |
[19] Nipponorthezinella guadalcanalia Last Modified in 12/05/13 | 8 | 61 | 61 | 0 | 0 | 0 | 0 | 1 |
[20] Arctorthezia occidentalis Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[21] Arctorthezia pseudoccidentalis Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[22] Arctorthezia cataphracta Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[23] Graminorthezia graminis Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[24] Graminorthezia tillandsiae Last Modified in 12/05/13 | 8 | 61 | 61 | 0 | 0 | 0 | 0 | 1 |
[25] Graminorthezia minor Last Modified in 12/05/13 | 13 | 56 | 56 | 0 | 0 | 0 | 0 | 1 |
[26] Orthezia urticae Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[27] Orthezia ambrosicola Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[28] Orthezia annae Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[29] Orthezia graminicola Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[30] Orthezia newcomeri Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[31] Praelongorthezia artemisiae Last Modified in 12/05/13 | 3 | 66 | 66 | 0 | 0 | 0 | 0 | 1 |
[32] Praelongorthezia praelonga Last Modified in 12/05/13 | 0 | 69 | 69 | 0 | 0 | 0 | 0 | 1 |
[33] Matilorthezia angolaensis Last Modified in 12/05/13 | 1 | 67 | 67 | 0 | 1 | 0 | 0 | 1 |
[34] Insignorthezia insignis Last Modified in 12/05/13 | 0 | 68 | 68 | 0 | 1 | 0 | 0 | 1 |
[35] Ortheziolacoccus madecassa Last Modified in 12/05/13 | 6 | 63 | 63 | 0 | 0 | 0 | 0 | 1 |
[36] Ortheziolamameti guineensis Last Modified in 12/05/13 | 3 | 66 | 66 | 0 | 0 | 0 | 0 | 1 |
[37] Ortheziola vejdovskyi Last Modified in 12/05/13 | 1 | 68 | 68 | 0 | 0 | 0 | 0 | 1 |
[38] Mixorthexia reynei Last Modified in 12/05/13 | 9 | 60 | 60 | 0 | 0 | 0 | 0 | 1 |
[39] Mixorthezia monticola Last Modified in 12/05/13 | 4 | 65 | 65 | 0 | 0 | 0 | 0 | 1 |
[40] Acropygorthezia williamsi Last Modified in 12/05/13 | 1 | 57 | 57 | 0 | 11 | 0 | 0 | 1 |
[41] † Mixorthezia dominicana Last Modified in 12/05/13 | 26 | 43 | 43 | 0 | 0 | 0 | 0 | 1 |
[42] † Mixorthezia kozari2 Last Modified in 12/05/13 | 25 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[43] † Newsteadia succini Last Modified in 12/05/13 | 25 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[44] † Arctorthezia antiqua Last Modified in 12/05/13 | 24 | 45 | 45 | 0 | 0 | 0 | 0 | 1 |
[45] † Arctorthezia baltica Last Modified in 12/05/13 | 25 | 44 | 44 | 0 | 0 | 0 | 0 | 1 |
[46] † Burmorthezia kotejai Last Modified in 12/05/13 | 24 | 43 | 43 | 0 | 2 | 0 | 0 | 1 |
[47] † Burmorthezia insolita Last Modified in 12/05/13 | 25 | 42 | 42 | 0 | 2 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 33344 | |
Project overview | 3512 | |
Media views | 6558 | Media search (5490 views); M102278 (1068 views); |
Matrix views | 2729 | Matrix landing page (2376 views); Female characters of Ortheziidae (353 views); |
Specimen list | 7857 | |
Taxon list | 12683 | |
Documents list | 5 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 391 | |
Media downloads | 1 | M102278 (1 download); |
Matrix downloads | 27 | Female characters of Ortheziidae (27 downloads); |
Project downloads | 363 |