Project 581: A. H. Turner, J. J. W. Sertich. 2010. Phylogenetic History of Simosuchus clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology. 30 (sp1):177-236.
Abstract
The bizarre crocodyliform Simosuchus clarki shares several derived cranial characteristics with basal notosuchian taxa. In the original phylogenetic analysis assessing placement of Simosuchus among other crocodyliforms, it was recovered as a basal notosuchian. Additional preparation of the holotype skull and postcranium, and discovery of additional specimens, provides new information on the suite of morphologies in this notosuchian. Morphological analysis of all available material of S. clarki yields 60 autapomorphies distributed across the skeleton. To provide an updated test of the phylogeny of Notosuchia and placement of Simosuchus within the clade, we added seven new characters and expanded taxon sampling to include recently described notosuchians. A phylogenetic analysis of 301 characters and 84 crocodyliforms supports the position of Simosuchus as sister to Libycosuchus, more derived than Araripesuchus and Uruguaysuchus, and basal to other ziphosuchian notosuchians. We did not recover an Anatosuchus + Simosuchus clade, nor a monophyletic Araripesuchus, but did recover a peirosaurid clade including Peirosaurus, Lomasuchus, Uberabasuchus, and Hamadasuchus (but not Stolokrosaurus). We also found weak support for inclusion of Anatosuchus, Araripesuchus, Peirosauridae, and Mahajangasuchidae within Notosuchia and therefore for the conclusion that all basal mesoeucrocodylians from Gondwana, with the exception of Stolokrosuchus, are notosuchians. The results of this analysis demonstrate that basal mesoeucrocodylian relationships are currently in a state of flux and the need for caution in naming higher-level taxa until a more stable consensus topology is achieved. Similarly, large gaps in the notosuchian record render inconclusive the biogeographic origins of Simosuchus and the clade it represents on Madagascar.Read the article »
Project DOI: 10.7934/P581, http://dx.doi.org/10.7934/P581
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 15747 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 301 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 696 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:301/0 |
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MorphoBank Project 581
MorphoBank Project 581
- Creation Date:
30 August 2011 - Publication Date:
18 November 2011 - Project views: 45668
- Matrix downloads: 30
This research
supported by
Authors' Institutions
- Denver Museum of Nature and Science
- Stony Brook University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
MorphoBank Curator Project Administrator | 1 | 1 | 1 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 | ||||
Maureen Admin Full membership | 85 | 0 | 0 | 301 | 0 | 0 | 15747 (15744, 0, 3) | 0 | 0 | 0 | ||||
Joseph Sertich Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M1433' (85 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] Gracilisuchus Last Modified in 12/05/13 | 108 | 193 | 193 | 0 | 0 | 0 | 0 | 3 |
[2] Terrestrisuchus Last Modified in 12/05/13 | 151 | 151 | 150 | 0 | 0 | 0 | 0 | 3 |
[3] Dibothrosuchus Last Modified in 12/05/13 | 77 | 224 | 224 | 0 | 0 | 0 | 0 | 3 |
[4] Protosuchus Last Modified in 12/05/13 | 47 | 259 | 254 | 0 | 0 | 0 | 0 | 3 |
[5] Hemiprotosuchus Last Modified in 12/05/13 | 175 | 126 | 126 | 0 | 0 | 0 | 0 | 3 |
[6] Orthosuchus Last Modified in 12/05/13 | 80 | 222 | 221 | 0 | 0 | 0 | 0 | 3 |
[7] Kayenta form Last Modified in 12/05/13 | 137 | 165 | 164 | 0 | 0 | 0 | 0 | 3 |
[8] Edentosuchus Last Modified in 12/05/13 | 180 | 127 | 121 | 0 | 0 | 0 | 0 | 3 |
[9] Zaraasuchus Last Modified in 12/05/13 | 224 | 80 | 77 | 0 | 0 | 0 | 0 | 3 |
[10] Gobiosuchus Last Modified in 12/05/13 | 82 | 224 | 219 | 0 | 0 | 0 | 0 | 3 |
[11] Sichuanosuchus Last Modified in 12/05/13 | 131 | 174 | 170 | 0 | 0 | 0 | 0 | 3 |
[12] Shantungosuchus Last Modified in 12/05/13 | 195 | 108 | 106 | 0 | 0 | 0 | 0 | 3 |
[13] Zosuchus Last Modified in 12/05/13 | 106 | 199 | 195 | 0 | 0 | 0 | 0 | 3 |
[14] Fruita form Last Modified in 12/05/13 | 118 | 184 | 183 | 0 | 0 | 0 | 0 | 3 |
[15] Notosuchus Last Modified in 12/05/13 | 24 | 281 | 277 | 0 | 0 | 0 | 0 | 3 |
[16] Comahuesuchus Last Modified in 12/05/13 | 155 | 150 | 146 | 0 | 0 | 0 | 0 | 3 |
[17] Mariliasuchus Last Modified in 12/05/13 | 106 | 199 | 195 | 0 | 0 | 0 | 0 | 3 |
[18] Uruguaysuchus Last Modified in 12/05/13 | 158 | 147 | 143 | 0 | 0 | 0 | 0 | 3 |
[19] Chimaeresuchus Last Modified in 12/05/13 | 206 | 96 | 95 | 0 | 0 | 0 | 0 | 3 |
[20] Malawisuchus Last Modified in 12/05/13 | 77 | 230 | 224 | 0 | 0 | 0 | 0 | 3 |
[21] Candidodon holo Last Modified in 12/05/13 | 295 | 6 | 6 | 0 | 0 | 0 | 0 | 3 |
[22] Simosuchus clarki Last Modified in 12/05/13 | 15 | 292 | 285 | 0 | 1 | 0 | 0 | 3 |
[23] Yacarerani boliviensis Last Modified in 12/05/13 | 94 | 209 | 207 | 0 | 0 | 0 | 0 | 3 |
[24] Sphagesaurus hueni Last Modified in 12/05/13 | 152 | 149 | 149 | 0 | 0 | 0 | 0 | 3 |
[25] Sphagesaurus montealtensis Last Modified in 12/05/13 | 105 | 198 | 196 | 0 | 0 | 0 | 0 | 3 |
[26] Adamantinasuchus Last Modified in 12/05/13 | 208 | 93 | 93 | 0 | 0 | 0 | 0 | 3 |
[27] Armadillosuchus arrudai Last Modified in 12/05/13 | 195 | 107 | 106 | 0 | 0 | 0 | 0 | 3 |
[28] Bretesuchus Last Modified in 12/05/13 | 196 | 110 | 105 | 0 | 0 | 0 | 0 | 3 |
[29] Baurusuchus Last Modified in 12/05/13 | 88 | 215 | 213 | 0 | 0 | 0 | 0 | 3 |
[30] Iberosuchus Last Modified in 12/05/13 | 112 | 197 | 189 | 0 | 0 | 0 | 0 | 3 |
[31] Libycosuchus Last Modified in 12/05/13 | 124 | 183 | 177 | 0 | 0 | 0 | 0 | 3 |
[32] Anatosuchus minor Last Modified in 12/05/13 | 45 | 259 | 256 | 0 | 0 | 0 | 0 | 3 |
[33] Araripesuchus gomesii Last Modified in 12/05/13 | 17 | 291 | 284 | 0 | 0 | 0 | 0 | 3 |
[34] Araripesuchus patagonicus Last Modified in 12/05/13 | 78 | 228 | 223 | 0 | 0 | 0 | 0 | 3 |
[35] Araripesuchus buitreraensi Last Modified in 12/05/13 | 184 | 125 | 117 | 0 | 0 | 0 | 0 | 3 |
[36] Araripesuchus wegeneri Last Modified in 12/05/13 | 28 | 275 | 272 | 0 | 1 | 0 | 0 | 3 |
[37] Araripesuchus tsangatsanga Last Modified in 12/05/13 | 37 | 268 | 263 | 0 | 1 | 0 | 0 | 3 |
[38] Lomasuchus Last Modified in 12/05/13 | 117 | 186 | 184 | 0 | 0 | 0 | 0 | 3 |
[39] Peirosaurus Last Modified in 12/05/13 | 197 | 107 | 104 | 0 | 0 | 0 | 0 | 3 |
[40] Hamadasuchus rebouli Last Modified in 12/05/13 | 78 | 223 | 223 | 0 | 0 | 0 | 0 | 3 |
[41] Uberabasuchus terrificus Last Modified in 12/05/13 | 142 | 163 | 159 | 0 | 0 | 0 | 0 | 3 |
[42] Montealtosuchus Last Modified in 12/05/13 | 73 | 228 | 228 | 0 | 0 | 0 | 0 | 3 |
[43] Theriosuchus Last Modified in 12/05/13 | 71 | 235 | 230 | 0 | 0 | 0 | 0 | 3 |
[44] Alligatorium Last Modified in 12/05/13 | 231 | 70 | 70 | 0 | 0 | 0 | 0 | 3 |
[45] Goniopholis simus Last Modified in 12/05/13 | 67 | 237 | 234 | 0 | 0 | 0 | 0 | 3 |
[46] Goniopholis stovalli Last Modified in 12/05/13 | 124 | 177 | 177 | 0 | 0 | 0 | 0 | 3 |
[47] Eutretanonew Last Modified in 12/05/13 | 116 | 187 | 185 | 0 | 0 | 0 | 0 | 3 |
[48] Calsoyasuchus Last Modified in 12/05/13 | 151 | 151 | 150 | 0 | 0 | 0 | 0 | 3 |
[49] Sunosuchus Last Modified in 12/05/13 | 185 | 116 | 116 | 0 | 0 | 0 | 0 | 3 |
[50] Shamosuchus Last Modified in 12/05/13 | 72 | 237 | 229 | 0 | 0 | 0 | 0 | 3 |
[51] Bernissartia Last Modified in 12/05/13 | 118 | 183 | 183 | 0 | 0 | 0 | 0 | 3 |
[52] Hylaeochampsa Last Modified in 12/05/13 | 231 | 70 | 70 | 0 | 0 | 0 | 0 | 3 |
[53] Allodaposuchus Last Modified in 12/05/13 | 177 | 127 | 124 | 0 | 0 | 0 | 0 | 3 |
[54] Glen rose form Last Modified in 12/05/13 | 112 | 189 | 189 | 0 | 0 | 0 | 0 | 3 |
[55] Borealosuchus Last Modified in 12/05/13 | 44 | 258 | 257 | 0 | 0 | 0 | 0 | 3 |
[56] Pristichampsus Last Modified in 12/05/13 | 74 | 229 | 227 | 0 | 0 | 0 | 0 | 3 |
[57] Eothoracosaurus Last Modified in 12/05/13 | 98 | 209 | 203 | 0 | 0 | 0 | 0 | 3 |
[58] Gavialis Last Modified in 12/05/13 | 23 | 281 | 278 | 0 | 0 | 0 | 0 | 3 |
[59] † Leidyosuchus canadensis Last Modified in 12/05/13 | 68 | 235 | 233 | 0 | 0 | 0 | 0 | 3 |
[60] Asiatosuchus germanicus Last Modified in 12/05/13 | 88 | 214 | 213 | 0 | 0 | 0 | 0 | 3 |
[61] Crocodylus Last Modified in 12/05/13 | 18 | 284 | 283 | 0 | 0 | 0 | 0 | 3 |
[62] Diplocynodon hantoniensis Last Modified in 12/05/13 | 41 | 261 | 260 | 0 | 0 | 0 | 0 | 3 |
[63] Alligator Last Modified in 12/05/13 | 17 | 285 | 284 | 0 | 0 | 0 | 0 | 3 |
[64] Hsisosuchus Last Modified in 12/05/13 | 112 | 194 | 189 | 0 | 0 | 0 | 0 | 3 |
[65] Pelagosaurus Last Modified in 12/05/13 | 59 | 247 | 242 | 0 | 0 | 0 | 0 | 3 |
[66] Steneosaurus Last Modified in 12/05/13 | 46 | 264 | 255 | 0 | 0 | 0 | 0 | 3 |
[67] Metriorhynchus superciliosus Last Modified in 12/05/13 | 80 | 223 | 221 | 0 | 0 | 0 | 0 | 3 |
[68] Metriorhynchus casamequel Last Modified in 12/05/13 | 187 | 114 | 114 | 0 | 0 | 0 | 0 | 3 |
[69] Geosaurus arauc Last Modified in 12/05/13 | 104 | 200 | 197 | 0 | 0 | 0 | 0 | 3 |
[70] Geosaurus suevi Last Modified in 12/05/13 | 183 | 120 | 118 | 0 | 0 | 0 | 0 | 3 |
[71] Dakosaurus maximus Last Modified in 12/05/13 | 223 | 78 | 78 | 0 | 0 | 0 | 0 | 3 |
[72] † Dakosaurus andiniensis Last Modified in 12/05/13 | 142 | 161 | 159 | 0 | 0 | 0 | 0 | 3 |
[73] Rhabdognathus m Last Modified in 12/05/13 | 129 | 175 | 172 | 0 | 0 | 0 | 0 | 3 |
[74] Sokotosuchus Last Modified in 12/05/13 | 232 | 69 | 69 | 0 | 0 | 0 | 0 | 3 |
[75] Dyrosaurus Last Modified in 12/05/13 | 123 | 178 | 178 | 0 | 0 | 0 | 0 | 3 |
[76] Hyposaurus Last Modified in 12/05/13 | 192 | 109 | 109 | 0 | 0 | 0 | 0 | 3 |
[77] Pholidosaurus Last Modified in 12/05/13 | 163 | 139 | 138 | 0 | 0 | 0 | 0 | 3 |
[78] Sarcosuchus Last Modified in 12/05/13 | 65 | 243 | 236 | 0 | 0 | 0 | 0 | 3 |
[79] Terminonaris robusta Last Modified in 12/05/13 | 92 | 215 | 209 | 0 | 0 | 0 | 0 | 3 |
[80] Pachycheilosuchus Last Modified in 12/05/13 | 210 | 94 | 91 | 0 | 0 | 0 | 0 | 3 |
[81] Rugosuchus nonganensis Last Modified in 12/05/13 | 129 | 176 | 172 | 0 | 0 | 0 | 0 | 3 |
[82] Mahajangasuchus Last Modified in 12/05/13 | 40 | 263 | 261 | 0 | 0 | 0 | 0 | 3 |
[83] Kaprosuchus saharicus Last Modified in 12/05/13 | 79 | 223 | 222 | 0 | 0 | 0 | 0 | 3 |
[84] Stolokrosuchus Last Modified in 12/05/13 | 88 | 215 | 213 | 0 | 0 | 0 | 0 | 3 |
[85] Argochampsa Last Modified in 12/05/13 | 145 | 158 | 156 | 0 | 0 | 0 | 0 | 3 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 45668 | |
Project overview | 3837 | |
Media views | 6746 | Media search (5558 views); M95901 (1188 views); |
Bibliography | 1251 | |
Matrix views | 2773 | Matrix landing page (2346 views); Simosuchus matrix (427 views); |
Specimen list | 3288 | |
Taxon list | 25877 | |
Views for media list | 1876 | |
Documents list | 20 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 366 | |
Project downloads | 336 | |
Matrix downloads | 30 | Simosuchus matrix (30 downloads); |