Project 568: J. C. Lamsdell, İ. Hoşgör, P. A. Selden. 2013. A New Ordovician Eurypterid (Arthropoda: Chelicerata) from South East Turkey: Evidence for a Cryptic Ordovician Record of Eurypterida. Gondwana Research. 23:354–366.
Abstract
A new species of eurypterid, Paraeurypterus anatoliensis gen. et sp. nov., is described from the Upper Ordovician (Katian) Şort Tepe Formation of southeast Turkey. The single specimen, preserving the carapace, mesosoma and fragments of appendages, appears morphologically intermediate between the eurypteroid families Dolichopteridae and Eurypteridae. P. anatoliensis retains the plesiomorphic conditions of crescentic eyes with enlarged palpebral lobes and a quadrate carapace with ornamentation consisting of small pustules but also displaying the derived characteristics of genal facets and a row of large acicular scales across the posterior of each tergite. Phylogenetic analysis incorporating each of the major eurypterine clades and all Eurypterina having a three-segmented genital operculum (the triploperculate condition) resolves eurypteroids to be an unnatural group, with Dolichopteridae and Eurypteridae forming part of a grade leading to diploperculate Eurypterina. P. anatoliensis is intermediate between the two eurypteroid families, as is ‘Eurypterus’ minor from the Pentland Hills of Scotland, which is shown to be a distinct genus and assigned to Pentlandopterus gen. nov. Using the phylogenetic topology to infer ghost ranges for each of the major eurypterid clades reveals that the majority of eurypterid superfamilies must have originated by the Katian, indicating a largely unsampled record of Ordovician eurypterids. The occurrence of poor dispersers such as Paraeurypterus in the Ordovician of Gondwana is puzzling, and it has been suggested that they dispersed to the continent during periods of sea level lowstand in the Sandbian and Hirnantian, however this does not explain the lack of Ordovician species in North America and Europe, given the well-sampled nature of these continents, and an alternative is proposed whereby eurypterids originated in Gondwana and radiated out to Laurentia and Baltica in the late Ordovician and early Silurian, thus explaining their sudden appearance in the European and North American rock record.Read the article »
Project DOI: 10.7934/P568, http://dx.doi.org/10.7934/P568
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 2970 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 81 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 195 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:81/0 |
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MorphoBank Project 568
MorphoBank Project 568
- Creation Date:
06 August 2011 - Publication Date:
25 November 2012 - Project views: 37462
- Media downloads: 2
- Matrix downloads: 10
Authors' Institutions
- Natural History Museum, London
- Capital Normal University
- University of Kansas
- TransAtlantic Petroleum Ltd.-Viking Int. Ltd.
Members
member name | taxa | specimens | media | media notes | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||||||||||
James Lamsdell Project Administrator | 45 | 1 | 1 | 0 | 81 | 0 | 0 | 0 | 0 | 2970 (2842, 0, 128) | 0 | 0 | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M1387' (45 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Weinbergina opitzi Last Modified in 12/05/13 | 1 | 67 | 67 | 0 | 13 | 0 | 0 | 1 |
[2] † Brachyopterus stubblefieldi Last Modified in 12/05/13 | 13 | 60 | 60 | 0 | 8 | 0 | 0 | 1 |
[3] † Parastylonurus ornatus Last Modified in 12/05/13 | 1 | 75 | 75 | 0 | 5 | 0 | 0 | 1 |
[4] † Stoermeropterus conicus Last Modified in 12/05/13 | 0 | 79 | 79 | 0 | 2 | 0 | 0 | 1 |
[5] † Stoermeropterus latus Last Modified in 12/05/13 | 14 | 65 | 65 | 0 | 2 | 0 | 0 | 1 |
[6] † Stoermeropterus nodosus Last Modified in 12/05/13 | 36 | 44 | 44 | 0 | 1 | 0 | 0 | 1 |
[7] † Vinetopterus struvei Last Modified in 12/05/13 | 4 | 75 | 75 | 0 | 2 | 0 | 0 | 1 |
[8] † Vinetopterus martini Last Modified in 12/05/13 | 17 | 62 | 62 | 0 | 2 | 0 | 0 | 1 |
[9] † Moselopterus ancylotelson Last Modified in 12/05/13 | 1 | 78 | 78 | 0 | 2 | 0 | 0 | 1 |
[10] † Moselopterus elongatus Last Modified in 12/05/13 | 4 | 75 | 75 | 0 | 2 | 0 | 0 | 1 |
[11] † Onychopterella augusti Last Modified in 12/05/13 | 15 | 63 | 63 | 0 | 3 | 0 | 0 | 1 |
[12] † Onychopterella kokomoensis Last Modified in 12/05/13 | 5 | 73 | 73 | 0 | 3 | 0 | 0 | 1 |
[13] † Dolichopterus macrocheirus Last Modified in 12/05/13 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 1 |
[14] † Eurypterus remipes Last Modified in 12/05/13 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 1 |
[15] † Hughmilleria socialis Last Modified in 12/05/13 | 2 | 76 | 76 | 0 | 3 | 0 | 0 | 1 |
[16] † Tylopterella boylei Last Modified in 12/05/13 | 46 | 34 | 34 | 0 | 1 | 0 | 0 | 1 |
[17] † Megalograptus ohioensis Last Modified in 12/05/13 | 0 | 75 | 75 | 0 | 6 | 0 | 0 | 1 |
[18] † Mixopterus kiaeri Last Modified in 12/05/13 | 3 | 76 | 76 | 0 | 2 | 0 | 0 | 1 |
[19] † Strobilopterus princetonii Last Modified in 12/05/13 | 12 | 68 | 68 | 0 | 1 | 0 | 0 | 1 |
[20] † Erieopterus microphthalmus Last Modified in 12/05/13 | 1 | 79 | 79 | 0 | 1 | 0 | 0 | 1 |
[21] † Rhenopterus diensti Last Modified in 12/05/13 | 0 | 75 | 75 | 0 | 6 | 0 | 0 | 1 |
[22] † Octoberaspis ushakovi Last Modified in 12/05/13 | 16 | 57 | 57 | 0 | 8 | 0 | 0 | 1 |
[23] † Loganamaraspis dunlopi Last Modified in 12/05/13 | 31 | 44 | 44 | 0 | 6 | 0 | 0 | 1 |
[24] † Diploaspis casteri Last Modified in 12/05/13 | 18 | 56 | 56 | 0 | 7 | 0 | 0 | 1 |
[25] † Paraeurypterus anatoliensis Last Modified in 12/05/13 | 43 | 38 | 38 | 0 | 0 | 0 | 0 | 1 |
[26] † Pentlandopterus minor Last Modified in 12/05/13 | 41 | 40 | 40 | 0 | 0 | 0 | 0 | 1 |
[27] † Chasmataspis laurencii Last Modified in 12/05/13 | 21 | 49 | 49 | 0 | 11 | 0 | 0 | 1 |
[28] † Buffalopterus pustulosus Last Modified in 12/05/13 | 34 | 45 | 45 | 0 | 2 | 0 | 0 | 1 |
[29] † Syntomopterella richardsoni Last Modified in 12/05/13 | 47 | 33 | 33 | 0 | 1 | 0 | 0 | 1 |
[30] † Dolichopterus jewetti Last Modified in 12/05/13 | 4 | 76 | 76 | 0 | 1 | 0 | 0 | 1 |
[31] † Ruedemannipterus stylonuroides Last Modified in 12/05/13 | 38 | 42 | 42 | 0 | 1 | 0 | 0 | 1 |
[32] † Clarkeipterus testudineus Last Modified in 12/05/13 | 46 | 34 | 34 | 0 | 1 | 0 | 0 | 1 |
[33] † Clarkeipterus otisius Last Modified in 12/05/13 | 67 | 13 | 13 | 0 | 1 | 0 | 0 | 1 |
[34] † Erieopterus eriensis Last Modified in 12/05/13 | 10 | 71 | 71 | 0 | 0 | 0 | 0 | 1 |
[35] † Eurypterus henkeni Last Modified in 12/05/13 | 9 | 71 | 71 | 0 | 1 | 0 | 0 | 1 |
[36] † Eurypterus tetragonophalmus Last Modified in 12/05/13 | 0 | 80 | 80 | 0 | 1 | 0 | 0 | 1 |
[37] † Eurypterus leopoldi Last Modified in 12/05/13 | 36 | 45 | 45 | 0 | 0 | 0 | 0 | 1 |
[38] † Eurypterus pittsfordensis Last Modified in 12/05/13 | 8 | 73 | 73 | 0 | 0 | 0 | 0 | 1 |
[39] † Eurypterus henningsmoeni Last Modified in 12/05/13 | 11 | 69 | 69 | 0 | 1 | 0 | 0 | 1 |
[40] † Orcanopterus manitoulinensis Last Modified in 12/05/13 | 13 | 66 | 66 | 0 | 2 | 0 | 0 | 1 |
[41] † Pterygotus anglicus Last Modified in 12/05/13 | 1 | 74 | 74 | 0 | 6 | 0 | 0 | 1 |
[42] † Adelophthalmus sievertsi Last Modified in 12/05/13 | 1 | 75 | 75 | 0 | 5 | 0 | 0 | 1 |
[43] † Eurypterus dekayi Last Modified in 12/05/13 | 1 | 79 | 79 | 0 | 1 | 0 | 0 | 1 |
[44] † Carcinosoma newlini Last Modified in 12/05/13 | 3 | 75 | 75 | 0 | 3 | 0 | 0 | 1 |
[45] † Nanahughmilleria norvegica Last Modified in 12/05/13 | 1 | 78 | 78 | 0 | 2 | 0 | 0 | 1 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 37462 | |
Project overview | 4114 | |
Matrix views | 3017 | Matrix landing page (2618 views); Turkish euypterid (399 views); |
Media views | 6708 | Media search (5767 views); M94106 (941 views); |
Specimen list | 3424 | |
Taxon list | 20197 | |
Documents list | 2 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 377 | |
Project downloads | 365 | |
Matrix downloads | 10 | Turkish euypterid (10 downloads); |
Media downloads | 2 | M94106 (2 downloads); |