Project 1200: Alan Turner. 2015. A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia. PLOS ONE. 10 (2):e0118116.
Abstract
The crocodyliform Shamosuchus is known from numerous Late Cretaceous localities in southern and eastern Mongolia and fragmentary remains from Uzbekistan. Seven species of Shamosuchus have been named from six localities in Mongolia and three in Uzbekistan. Six species originally described as Paralligator were later referred to Shamosuchus. Only the type species, Shamosuchus djadochtaensis has been examined in detail. Many of the named species of Shamosuchus show striking similarity in size and cranial morphology but most are based on partial remains suggesting that the true species diversity is overestimated.A review of all species referred to Shamosuchus recognizes three valid taxa: Shamosuchus djadochtaensis, S. gradilifrons, and S. major. Shamosuchus sungaricus, S. borealis, and S. karakalpakensis are nomena dubia, whereas S. ancestralis, S. ulgicus, S. tersus, and S. ulanicus are junior synonyms of S. gradilifrons. Phylogenetic analysis of 318 phenotypic characters recovers a Paralligatoridae clade consisting of Shamosuchus, Rugosuchus, Batrachomimus, Glen Rose Form, and Wannchampsus. Shamosuchus is non-monophyletic: S. djadochatensis is near the base of Paralligatoridae whereas S. gradilifrons + S. major are the most deeply nested. The name Paralligator is resurrected for this clade. Rugosuchus and Batrachomimus are sister taxa to Paralligator. Paralligatoridae is closely related to Theriosuchus, hylaeochampsids and a speciose Allodaposuchus clade, which together are the sister group of Borealosuchus plus Crocodylia. These results support the presence of a diverse clade in eastern Asia and western North America throughout the Cretaceous with origins in the Late Jurassic.
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Article DOI: 10.1371/journal.pone.0118116
Project DOI: 10.7934/P1200, http://dx.doi.org/10.7934/P1200
This project contains | Matrices |
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Download Project SDD File | Total scored cells: 21071 Total media associated with cells: 0 Total labels associated with cell media: 0 |
Characters | |
Total characters: 321 Total characters with associated media: 0 Total characters with media with labels: 0 Total character states: 731 Total character states with associated media: 0 Total character states with media with labels:0 Total unordered/ordered characters:321/0 |
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MorphoBank Project 1200
MorphoBank Project 1200
- Creation Date:
03 June 2014 - Publication Date:
18 February 2015 - Project views: 44271
- Media downloads: 3
- Matrix downloads: 146
This research
supported by
Authors' Institutions
- Stony Brook University
Members
member name | taxa | specimens | media | chars | character
| cell scorings (scored, NPA, "-") | cell
| rules | ||||||
Alan Turner Project Administrator | 120 | 1 | 1 | 321 | 0 | 0 | 21071 (20840, 0, 231) | 0 | 0 | 0 | ||||
MorphoBank Curator Full membership | 0 | 0 | 0 | 0 | 0 | 0 | 0 (0, 0, 0) | 0 | 0 | 0 |
Taxonomic Overview for Matrix 'M2711' (109 Taxa)
taxon | unscored cells |
scored cells |
no cell support |
NPA cells |
"-" cells | cell images | labels on cell images |
member access |
[1] † Gracilisuchus stipanicicorum Last Modified in 02/19/15 | 114 | 202 | 0 | 0 | 5 | 0 | 0 | 2 |
[2] † Terrestrisuchus gracilis Last Modified in 02/19/15 | 157 | 159 | 0 | 0 | 6 | 0 | 0 | 2 |
[3] † Dibothrosuchus elaphros Last Modified in 02/19/15 | 79 | 236 | 0 | 0 | 6 | 0 | 0 | 2 |
[4] † Protosuchus richardsoni Last Modified in 02/19/15 | 50 | 273 | 0 | 0 | 4 | 0 | 0 | 2 |
[5] † Hemiprotosuchus leali Last Modified in 02/19/15 | 194 | 126 | 0 | 0 | 1 | 0 | 0 | 2 |
[6] † Orthosuchus stormbergi Last Modified in 02/19/15 | 91 | 226 | 0 | 0 | 5 | 0 | 0 | 2 |
[7] † Kayenta form Last Modified in 02/19/15 | 142 | 174 | 0 | 0 | 6 | 0 | 0 | 2 |
[8] † Edentosuchus tienshanensis Last Modified in 02/19/15 | 193 | 134 | 0 | 0 | 0 | 0 | 0 | 2 |
[9] † Zaraasuchus shepardi Last Modified in 02/19/15 | 242 | 82 | 0 | 0 | 0 | 0 | 0 | 2 |
[10] † Gobiosuchus kielanae Last Modified in 02/19/15 | 88 | 235 | 0 | 0 | 3 | 0 | 0 | 2 |
[11] † Sichuanosuchus shuhanensis Last Modified in 02/19/15 | 135 | 188 | 0 | 0 | 2 | 0 | 0 | 2 |
[12] † Shantungosuchus hangjinensis Last Modified in 02/19/15 | 208 | 113 | 0 | 0 | 2 | 0 | 0 | 2 |
[13] † Zosuchus davidsoni Last Modified in 02/19/15 | 109 | 215 | 0 | 0 | 1 | 0 | 0 | 2 |
[14] † Fruitachampsa callisoni Last Modified in 02/19/15 | 120 | 200 | 0 | 0 | 2 | 0 | 0 | 2 |
[15] † Notosuchus terrestris Last Modified in 02/19/15 | 25 | 298 | 0 | 0 | 2 | 0 | 0 | 2 |
[16] † Comahuesuchus brachybuccalis Last Modified in 02/19/15 | 156 | 167 | 0 | 0 | 2 | 0 | 0 | 2 |
[17] † Mariliasuchus amarali Last Modified in 02/19/15 | 108 | 215 | 0 | 0 | 2 | 0 | 0 | 2 |
[18] † Uruguaysuchus aznarezi Last Modified in 02/19/15 | 162 | 162 | 0 | 0 | 1 | 0 | 0 | 2 |
[19] † Chimaeresuchus paradoxus Last Modified in 02/19/15 | 218 | 103 | 0 | 0 | 1 | 0 | 0 | 2 |
[20] † Malawisuchus mwakasyungutiensis Last Modified in 02/19/15 | 80 | 246 | 0 | 0 | 1 | 0 | 0 | 2 |
[21] † Candidodon itapecuruense Last Modified in 02/19/15 | 301 | 19 | 0 | 0 | 1 | 0 | 0 | 2 |
[22] † Simosuchus clarki Last Modified in 02/19/15 | 16 | 311 | 0 | 0 | 1 | 0 | 0 | 2 |
[23] † Yacarerani boliviensis Last Modified in 02/19/15 | 95 | 226 | 0 | 0 | 2 | 0 | 0 | 2 |
[24] † Sphagesaurus hueni Last Modified in 02/19/15 | 159 | 160 | 0 | 0 | 2 | 0 | 0 | 2 |
[25] † Sphagesaurus montealtensis Last Modified in 02/19/15 | 105 | 216 | 0 | 0 | 2 | 0 | 0 | 2 |
[26] † Adamantinasuchus navae Last Modified in 02/19/15 | 223 | 98 | 0 | 0 | 0 | 0 | 0 | 2 |
[27] † Armadillosuchus arrudai Last Modified in 02/19/15 | 204 | 117 | 0 | 0 | 1 | 0 | 0 | 2 |
[28] † Bretesuchus bonapartei Last Modified in 02/19/15 | 206 | 119 | 0 | 0 | 1 | 0 | 0 | 2 |
[29] † Baurusuchus pachecoi Last Modified in 02/19/15 | 99 | 223 | 0 | 0 | 1 | 0 | 0 | 2 |
[30] † Iberosuchus macrodon Last Modified in 02/19/15 | 132 | 197 | 0 | 0 | 0 | 0 | 0 | 2 |
[31] † Libycosuchus brevirostris Last Modified in 02/19/15 | 132 | 195 | 0 | 0 | 0 | 0 | 0 | 2 |
[32] † Anatosuchus minor Last Modified in 02/19/15 | 48 | 275 | 0 | 0 | 1 | 0 | 0 | 2 |
[33] † Araripesuchus gomesii Last Modified in 02/19/15 | 19 | 308 | 0 | 0 | 1 | 0 | 0 | 2 |
[34] † Araripesuchus patagonicus Last Modified in 02/19/15 | 80 | 245 | 0 | 0 | 1 | 0 | 0 | 2 |
[35] † Araripesuchus buitreraensis Last Modified in 02/19/15 | 190 | 138 | 0 | 0 | 1 | 0 | 0 | 2 |
[36] † Araripesuchus wegeneri Last Modified in 02/19/15 | 29 | 293 | 0 | 0 | 2 | 0 | 0 | 2 |
[37] † Araripesuchus tsangatsangana Last Modified in 02/19/15 | 41 | 284 | 0 | 0 | 1 | 0 | 0 | 2 |
[38] † Lomasuchus palpebrosus Last Modified in 02/19/15 | 122 | 200 | 0 | 0 | 1 | 0 | 0 | 2 |
[39] † Peirosaurus torminni Last Modified in 02/19/15 | 207 | 115 | 0 | 0 | 2 | 0 | 0 | 2 |
[40] † Hamadasuchus rebouli Last Modified in 02/19/15 | 78 | 242 | 0 | 0 | 1 | 0 | 0 | 2 |
[41] † Uberabasuchus terrificus Last Modified in 02/19/15 | 151 | 174 | 0 | 0 | 0 | 0 | 0 | 2 |
[42] † Montealtosuchus arrudacamposi Last Modified in 02/19/15 | 73 | 246 | 0 | 0 | 2 | 0 | 0 | 2 |
[43] † Trematochampsa Last Modified in 02/19/15 | 217 | 108 | 0 | 0 | 0 | 0 | 0 | 2 |
[44] † Mahajangasuchus insignis Last Modified in 02/19/15 | 43 | 280 | 0 | 0 | 0 | 0 | 0 | 2 |
[45] † Kaprosuchus saharicus Last Modified in 02/19/15 | 79 | 243 | 0 | 0 | 0 | 0 | 0 | 2 |
[46] † Stolokrosuchus lapparenti Last Modified in 02/19/15 | 93 | 228 | 0 | 0 | 1 | 0 | 0 | 2 |
[47] † Hsisosuchus chungkingensis Last Modified in 02/19/15 | 121 | 202 | 0 | 0 | 3 | 0 | 0 | 2 |
[48] † Pelagosaurus typus Last Modified in 02/19/15 | 62 | 263 | 0 | 0 | 1 | 0 | 0 | 2 |
[49] † Steneosaurus bollensis Last Modified in 02/19/15 | 55 | 274 | 0 | 0 | 1 | 0 | 0 | 2 |
[50] † Metriorhynchus superciliosus Last Modified in 02/19/15 | 87 | 236 | 0 | 0 | 0 | 0 | 0 | 2 |
[51] † Metriorhynchus casamequelai Last Modified in 02/19/15 | 201 | 120 | 0 | 0 | 0 | 0 | 0 | 2 |
[52] † Cricosaurus araucanensis Last Modified in 02/19/15 | 109 | 214 | 0 | 0 | 1 | 0 | 0 | 2 |
[53] † Cricosaurus suevicus Last Modified in 02/19/15 | 199 | 124 | 0 | 0 | 0 | 0 | 0 | 2 |
[54] † Dakosaurus maximus Last Modified in 02/19/15 | 238 | 83 | 0 | 0 | 0 | 0 | 0 | 2 |
[55] † Dakosaurus andiniensis Last Modified in 02/19/15 | 148 | 174 | 0 | 0 | 1 | 0 | 0 | 2 |
[56] † Rhabdognathus Last Modified in 02/19/15 | 132 | 191 | 0 | 0 | 1 | 0 | 0 | 2 |
[57] † Sokotosuchus ianwilsoni Last Modified in 02/19/15 | 251 | 70 | 0 | 0 | 0 | 0 | 0 | 2 |
[58] † Dyrosaurus Last Modified in 02/19/15 | 126 | 194 | 0 | 0 | 1 | 0 | 0 | 2 |
[59] † Hyposaurus rogersii Last Modified in 02/19/15 | 203 | 118 | 0 | 0 | 0 | 0 | 0 | 2 |
[60] † Pholidosaurus purbeckensis Last Modified in 02/19/15 | 174 | 148 | 0 | 0 | 0 | 0 | 0 | 2 |
[61] † Sarcosuchus imperator Last Modified in 02/19/15 | 66 | 261 | 0 | 0 | 1 | 0 | 0 | 2 |
[62] † Terminonaris robusta Last Modified in 02/19/15 | 93 | 233 | 0 | 0 | 1 | 0 | 0 | 2 |
[63] † Elosuchus cherifiensis Last Modified in 02/19/15 | 217 | 103 | 0 | 0 | 1 | 0 | 0 | 2 |
[64] † Meridiosaurus vallisparadisi Last Modified in 02/19/15 | 264 | 56 | 0 | 0 | 1 | 0 | 0 | 2 |
[65] † Oceanosuchus boecensis Last Modified in 02/19/15 | 236 | 85 | 0 | 0 | 0 | 0 | 0 | 2 |
[66] † Paluxysuchus newmani Last Modified in 02/19/15 | 112 | 208 | 0 | 0 | 1 | 0 | 0 | 2 |
[67] † Goniopholis simus Last Modified in 02/19/15 | 68 | 255 | 0 | 0 | 1 | 0 | 0 | 2 |
[68] † Amphicotylus lucasii Last Modified in 02/19/15 | 123 | 196 | 0 | 0 | 2 | 0 | 0 | 2 |
[69] † Calsoyasuchus valliceps Last Modified in 02/19/15 | 161 | 159 | 0 | 0 | 2 | 0 | 0 | 2 |
[70] † Sunosuchus junggarensis Last Modified in 02/19/15 | 192 | 127 | 0 | 0 | 2 | 0 | 0 | 2 |
[71] EutreALL Last Modified in 06/03/14 | 50 | 261 | 0 | 0 | 10 | 0 | 0 | 2 |
[72] † Goniopholis baryglyphaeus Last Modified in 02/19/15 | 196 | 125 | 0 | 0 | 0 | 0 | 0 | 2 |
[73] † Alligatorium Last Modified in 02/19/15 | 244 | 77 | 0 | 0 | 0 | 0 | 0 | 2 |
[74] † Theriosuchus pusillus Last Modified in 02/19/15 | 59 | 264 | 0 | 0 | 3 | 0 | 0 | 2 |
[75] † Theriosuchus sympiestodon Last Modified in 02/19/15 | 242 | 75 | 0 | 0 | 4 | 0 | 0 | 2 |
[76] † Theriosuchus guimarotae Last Modified in 02/19/15 | 171 | 151 | 0 | 0 | 1 | 0 | 0 | 2 |
[77] † Wannchampsus kirpachi Last Modified in 02/19/15 | 88 | 232 | 0 | 0 | 1 | 0 | 0 | 2 |
[78] † Shamosuchus major Last Modified in 02/19/15 | 164 | 152 | 0 | 0 | 7 | 0 | 0 | 2 |
[79] † Shamosuchus djadochtaensis Last Modified in 02/19/15 | 76 | 252 | 0 | 0 | 1 | 0 | 0 | 2 |
[80] † Shamosuchus gradilifrons Last Modified in 02/19/15 | 40 | 278 | 0 | 0 | 8 | 0 | 0 | 2 |
[81] † Shamosuchus gradilifrons holotype Last Modified in 02/19/15 | 105 | 218 | 0 | 0 | 4 | 0 | 0 | 2 |
[82] † Shamosuchus tersus Last Modified in 02/19/15 | 138 | 182 | 0 | 0 | 4 | 0 | 0 | 2 |
[83] † Shamosuchus ancestralis Last Modified in 02/19/15 | 230 | 91 | 0 | 0 | 1 | 0 | 0 | 2 |
[84] † Shamosuchus ulanicus Last Modified in 02/19/15 | 89 | 228 | 0 | 0 | 9 | 0 | 0 | 2 |
[85] † Shamosuchus ulgicus Last Modified in 02/19/15 | 313 | 7 | 0 | 0 | 1 | 0 | 0 | 2 |
[86] † Rugosuchus nonganensis Last Modified in 02/19/15 | 133 | 191 | 0 | 0 | 1 | 0 | 0 | 2 |
[87] † Batrachomimus pastosbonensis Last Modified in 02/19/15 | 139 | 173 | 0 | 0 | 9 | 0 | 0 | 2 |
[88] Glen rose form Last Modified in 06/03/14 | 119 | 201 | 0 | 0 | 1 | 0 | 0 | 2 |
[89] † Bernissartia fagesii Last Modified in 02/19/15 | 125 | 195 | 0 | 0 | 1 | 0 | 0 | 2 |
[90] † Hylaeochampsa vectiana Last Modified in 02/19/15 | 234 | 86 | 0 | 0 | 1 | 0 | 0 | 2 |
[91] † Iharkutosuchus makadii Last Modified in 02/19/15 | 94 | 226 | 0 | 0 | 1 | 0 | 0 | 2 |
[92] † Acynodon iberoccitanus Last Modified in 02/19/15 | 95 | 208 | 0 | 0 | 18 | 0 | 0 | 2 |
[93] † Acynodon adriaticus Last Modified in 02/19/15 | 105 | 206 | 0 | 0 | 12 | 0 | 0 | 2 |
[94] † Allodaposuchus precedens Last Modified in 02/19/15 | 179 | 144 | 0 | 0 | 1 | 0 | 0 | 2 |
[95] † Allodaposuchus subjuniperus Last Modified in 02/19/15 | 130 | 190 | 0 | 0 | 1 | 0 | 0 | 2 |
[96] † Allodaposuchus cf. precedens Last Modified in 02/19/15 | 111 | 198 | 0 | 0 | 12 | 0 | 0 | 2 |
[97] † Pachycheilosuchus trinquei Last Modified in 02/19/15 | 226 | 98 | 0 | 0 | 0 | 0 | 0 | 2 |
[98] † Susisuchus anatoceps Last Modified in 02/19/15 | 157 | 164 | 0 | 0 | 0 | 0 | 0 | 2 |
[99] † Isisfordia duncani Last Modified in 02/19/15 | 44 | 267 | 0 | 0 | 11 | 0 | 0 | 2 |
[100] † Borealosuchus formidabilis Last Modified in 02/19/15 | 46 | 275 | 0 | 0 | 1 | 0 | 0 | 2 |
[101] † Argochampsa krebsi Last Modified in 02/19/15 | 147 | 175 | 0 | 0 | 1 | 0 | 0 | 2 |
[102] † Pristichampsus vorax Last Modified in 02/19/15 | 76 | 246 | 0 | 0 | 1 | 0 | 0 | 2 |
[103] † Eothoracosaurus mississippiensis Last Modified in 02/19/15 | 99 | 227 | 0 | 0 | 1 | 0 | 0 | 2 |
[104] Gavialis gangeticus Last Modified in 06/03/14 | 23 | 300 | 0 | 0 | 1 | 0 | 0 | 2 |
[105] † Leidyosuchus canadensis Last Modified in 02/19/15 | 68 | 254 | 0 | 0 | 1 | 0 | 0 | 2 |
[106] † Asiatosuchus germanicus Last Modified in 02/19/15 | 93 | 228 | 0 | 0 | 1 | 0 | 0 | 2 |
[107] Crocodylus niloticus Last Modified in 06/03/14 | 18 | 303 | 0 | 0 | 1 | 0 | 0 | 2 |
[108] † Diplocynodon hantonensis Last Modified in 02/19/15 | 41 | 280 | 0 | 0 | 1 | 0 | 0 | 2 |
[109] Alligator mississippiensis Last Modified in 06/03/14 | 17 | 304 | 0 | 0 | 1 | 0 | 0 | 2 |
Project views
type | number of views | Individual items viewed (where applicable) |
Total project views | 44271 | |
Project overview | 5150 | |
Media views | 6410 | Media search (5592 views); M333576 (818 views); |
Matrix views | 3545 | Matrix landing page (2981 views); A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia (564 views); |
Specimen list | 3161 | |
Taxon list | 22420 | |
Documents list | 2404 | |
Bibliography | 1181 |
Project downloads
type | number of downloads | Individual items downloaded (where applicable) |
Total downloads from project | 678 | |
Matrix downloads | 146 | A review of Shamosuchus and Paralligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia (146 downloads); |
Document downloads | 67 | MrBayes Results Treefile (10 downloads); MrBayes analysis file (9 downloads); TNT file (18 downloads); Isisfordia Primary Analysis TNT file (8 downloads); TNT constraint file (12 downloads); Incipient procoely Tree File (1 download); Derive palate and procoely TNT file (1 download); Derived palate TNT file (2 downloads); Derived procoely TNT file (2 downloads); Incipient procoely TNT file (3 downloads); Derived palate strict consensus tree file (1 download); |
Project downloads | 462 | |
Media downloads | 3 | M333576 (3 downloads); |