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Project 1117: I. J. Garzón-Orduña, O. Marini-Filho, S. G. Johnson, C. M. Penz. 2013. Phylogenetic relationships of Hamadryas (Nymphalidae: Biblidinae) based on the combined analysis of morphological and molecular data. Cladistics. 29 (6):629-642.
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